14 Art. 8. — S. Goto and Y. Matsudaira : 



and appear to open on the cuticle. The dermal glands before 

 noted as lying on either side of the oral sucker are probably special 

 developments of these subcuticular cell groups. As to the lymph 

 spaces they are especially well developed in this species, and 

 although we have not been able to make out their mutual connec- 

 tions thoroughly, there is no doubt that they consist in the 

 main of longitudinal vessels extending nearly the entire length of 

 the body. At the level of the intestinal cseca one can count 

 as many as fifteen spaces in a cross section, most of them im- 

 mediately surrounding the cseca but some lying at some distance 

 away from them. At the level of the genital pore they are 

 situated around the intestinal ca?ca and on the dorsal and 

 lateral sides of the penis sac; and at the two ends of the 

 body, where the spaces are less in number, they are situated 

 on the dorsal side of the respective suckers. In some sections it 

 appeared to us that some of these spaces fused and communicated 

 with the neighbouring ones. 



There is no doubt that this new genus is most nearly allied to 

 the Paramphistomatidœ. Aside from the general form of the body 

 with the two suckers at either end, the character of the prepharynx, 

 the development of a muscular pharynx in its course, and 

 above all the presence of a lymph system are characters which 

 can be noted in varying degrees in the members of that 

 family. Thus Fischoeder [p. 491] mentions a certain degree of 

 the winding of the prepharynx ('oesophagus' according to his 

 terminology) as a distinguishing character of the Paramphistomat- 

 idœ, but it is carried to an extreme in our new genus, as above 

 described. Again the presence of gland cells around the pre- 

 pharynx appears to be a characteristic of that family, so far 

 as its members have been studied with reference to this point. 

 A muscular pharynx, more or less well developed, somewhere 

 in the course of the prepharynx, is found in such forms as 

 Watso/iius watsoni, PseudodLscus haw/res Li, OpLsthodiscus diplodis- 

 ciLdes, SchlzamphLstomum spLnulosum and SchLzamph. scleropoiim. 

 As to the lymph system its significance has been so ably pointed 

 out not long ago by Odhner that I deem it sufficient merely to refer 



