Cope.] ^± [April 4, 



metacarpus, and nearly half as long as the outer (5th) metacarpal. It 

 articulates with a thick V-shaped facet of the cuneiform. Its extremity 

 is obtuse and expanded. The trapezium is large and attached to its 

 metacarpus laterally, sending a process downwards posteriorly. It sup- 

 ports a narrow articular surface for the metacarpus of a small pollex or 

 * inner digit, which is not preserved. The trapezoid is smaller and of a 

 triangular outline, with the base forwards. The magnum is a rather 

 small bone articulating as usual with the metatarsals 2 and 3. It is 

 depressed in front. The unciform is a large bone with a considerable 

 external anterior surface. Two-thirds of its upper surface is in contact 

 with the cuneiform, the remaining part projecting upwards with convex 

 face to unite with the lunare. Eelow it supports metatarsals 4 and 5. 



There were probably five digits of the fore foot, the inner small or 

 rudimental. The proportions are stouter than in the dogs, but not so 

 much so as in the bears. The phalanges have a length similar to that seen 

 in some bears, but the metatarsals are more elongate. The lengths of 

 the latter are, fifth shortest, then 2d, 3d and 4th. Their condyles are 

 broad, with median keel behind, and shallow supracondylar fossa in 

 front. The first phalanges are about one-third the length of the meta- 

 carpals ; the second of digit No. 2 broad and stout and half as long as 

 the phalange of the first row. An ungueal phalange has a singular 

 form, so that the claw might be supposed to have a subungulate charac- 

 acter. It is flat, considerably broader than high and with expanded and 

 obtuse extremity. The articular extremity is depressed and transverse 

 concave in vertical, convex in transverse section. The anterior three- 

 fifths of the superior middle line is occupied by a deep gaping fissure, 

 which separates the extremity into two points. The inferior face is 

 entirely fiat, there being no tendinous tuberosity. The sides are grooved, 

 and give entrance each to a large arterial foramen proximally. These 

 claws resemble those of Mesonyx, and differ remarkably from those of 

 existing terrestrial Garnivora. 



Of hinder limb the only characteristic pieces remaining are. the navicu- 

 lar, cuboid, and an external cuneiform bone. The cuboid is rather 

 stout, with a slight concave facet at one extremity and two at the other, 

 one of them smaller and sublateral. The navicular is wide and flat, and 

 with a strongly concave astragaline facet. Below, it presents two deep 

 oblique concave facets for the cuneiforms, with a small sublateral one 

 on the outer side. The facets of the cuboid and astragalus indicate four 

 well developed digits and another perhaps smaller one. Thus in this 

 genus they were on both limbs probably 5-5, with the inner small. 



The cranium is fractured above. There remain the squamosal and 

 periotic bones, occipital condyles, malar and part of maxillary, both 

 premaxillaries and the greater part of both mandibular rami. The 

 squamosal process of the zygoma is produced inferiorly far below T the 

 auditory meatus, even further than in the bears. Its proximal portion 

 includes, on the lower face, a strong groove at right angles to the axis 



