4 ART. :). T. ICHIMURA : ON THE FORMATION OF 



may lu; uUied 2)igüiciitc5. Zimniermanii'^ describes the "pigment- 

 secretion" as a certain componnd of antliocyan with some unknown 

 tannin sulistance. According to chemical examinations, however, 

 the violet crystals, which I have observed, proved to be nothing 

 else than those of antliocyan. 



In the following pages I shall describe the development of 

 anthocyiin in Japanese Hortense in four phases. In Kanazawa, 

 tiie ilowering period of this plant lasts about four months, beginn- 

 ing in June and ending in September. My observations were 

 carried on during the vears, 1900 and 1*J()I. 



11. Different Phases of the Development of the Petaloid Calyx. 



The Fir.«T Phase. 



Diu-ation of observation: June 1 — July 1. Exlcrixtl fonn. — 

 At the biginning of June most of the peripheral llower-buds 

 swell and open gradually. (Fig. 1.) Xow each young llower 

 nutates upon a curved stalk and already carries four or five 

 involved petaloid c;dyx-leaves. Among the latter, the one 

 which is the lowest in position is the largest and entirely 

 covers the other. At this stage, particularly in the largest calyx 

 leaf, the predominance of hyponastic growth can l)e traced, but, 

 later, epinastic growth becomes the more powerful. Soon after- 

 ward, each curved ilower stalk tends to stand erect and the 

 ilower itself to open. Even in full bloom, each calyx-leaf, remains 

 to the last phase without losing its spoon shape. 



5) Ziiuineniiaiiii, A. — Die Butanisclie Mikrottchnik. 1892, p. 104. 



