A Monograph of the Genus Alaria. 41 



dans toutes les parties de la lame" of A. ßstidosa^) The remark 

 by Setchell and Gaedner refers to these Unes. 



Güignard's observation on the mentioned species have all been 

 done on herbarium specimens. It is rather excusable that he has 

 mistaken the mucilage glands of A. fistulosa as the mucilage lacu- 

 nae, and overlooked the similar but smaller cells in the fronds of 

 Undaria pinnatlfida. His statement disproving the conclusion by 

 Kjellmax"^) that Alaria plants have no mucilage canals is therefore 

 to be re-disproved. 



In a fully matured frond of A. ochotensis Yendo the transition 

 region is so rich in the roundish glands as to turn the part of the 

 blade, when dried, into a dark reddish-brown colour. In the upper 

 part of the blade as well as in the membranous sterile part of the 

 sporophylls, the glands are much elongated, furcated and armed 

 with short lateral processes, and a few of them are often fused 

 together (Plate XIX, figs. 2-3). The mucilage lacunae of Laminât- ia 

 occur in some species, for example, Laminar ia angustata Kjellm., in 

 detached groups of incomplete network.^) The mucilage glands of 

 A. ochotensis approach in general aspect, though of course geneti- 

 cally differing from, the mucilage lacunae of L. angustata. It is 

 too complicated for a fucosan reservoir. 



The mucilage glands of the other species of Alaria have more 

 or less fixed character in each. The three examples described 

 above illustrate extreme cases of the primitive and well-developed 

 state of the glands. Under the descriptive part some accounts of 

 the gland of each species will be given, referring to the above 

 remarks. 



1) Gtjignard : Observations stir Appareil Mucilère des Laminariacèes, p. 43. 



2) Kjelljian : Handbok i Skandinaviens Hafsalgflora, p. 19. 



3) Miyabe: Laminaria Indnsfry of Hokkailo, Pla'e 29, fig. 5. 



