39 



anlerioily and doisally near the lip ol' Ihe basal joini of all the Garypidae Hans., 

 wiielher this joint is long or short, a long tissnre, which is directed obliquely for- 

 wards (cf. figs. 7-8 a, /■) (Fl. II, ligs. le, 7 f and 8 f). If the basal joints of f. insl. 

 (lariipiiuis nobilis n. sp. and Gaii/piis irrii(]alus Sim. were not homologous, Ihe 

 pieviously mentioned lissure should have had a dillerent position in the two 

 forms. The position of Börner's „llexor femoris 11" in Garijpim L. K. as well as 

 in Garypiniis Dad. tells the same story. 



If we are thus right in regarding the basal as well as the tibial femoral 

 parts as being homologous in all the Gnrypidae Hans., and if our assumption 

 of the homonolgy of all the trochantins and distal portions of the femurs in the 

 Cheliferinae Sim. is true, it will certainly be necessary with Borner to homologize 

 the trochantin of Chclifer Geof. with the basal femoral ]iart of all the other genera, 

 and the distal femoral part of that genus with the til)ial femoral part of these. 

 The corresponding position of the flexor-muscle of the femoral tibial part in the 

 Cheliferinae Sim. as well as in the Garypidae Hans, is in favour of this inter- 

 pretation. 



With regard to the differences between the femoral parts and the articulation 

 between them in Gnrypiis L. K. and Chthoniiis-Obisiiim C. K. I refer to Hansen (49. 

 p. 224), who has pointed out, that the tibial femoral part has a tendency to become 

 more and more resembling a tibia in shape as well as in articulation, going from 

 the former of these genera to the latter passing Ideobisiiim Balz, and Obisinm C. K. 

 The tibial part of Chthonius C. K. has really in almost all respects become a tibia. 



IV. Femur of Fourth Pair of Legs. 



The joints of the fourth pair of legs have not been interpreted in a difTerent 

 way by the authors, because their femora do not provide so striking variations, as 

 those of the first pair do. The femur of the fourth pair is in all genera, but one 

 Chiridiiim Menge, provided with a longer or shorter trochantin, set off I'rom the 

 rest of the femur by a strictum and without any muscles, at least in all genera 

 except Chthonius C. K. That the trochantin of the fourth pair of legs is homologous 

 in all the Chelonethi, may be regarded as self-evident. The (juestion remains, 

 whether the two portions of the femur of the fourth pair of legs correspond to 

 those of the first pair, or not. Such an identification seems at least to be very 

 probable'), when we remember, that all the other joints of the legs are "homotype" 

 to each other, that there seems to be a kind of specialized articulation between 

 the two femoi'al portions in Chthonius C. K., and that there is a reduction in the 

 length and development of the trochantin of the fourth pair, corresponding to that 

 of the basal femoral portion of the first pair, if we go from Chthonius C. K. to 

 Garypus-Clielifer. If one may be allowed to express oneself paradoxically, one can 

 say, that the femur of the fourth pair of legs in Chthonius C. K. stands on the 



') The remarkably well developed articulation between the trochantin and the femur proper of 

 the fourth pair of lefjs in Fcaella mirabilis Ell. is strongly in favour of this theory (cf. Kllingsen 72). 



