WHITE— STUDIES OF INHERITANCE IN PISUM. 511 



varieties having orange-red seed coats and obtained in F^ progeny 

 with seed coats varying from almost colorless ? to intense orange- 

 red — the variation in coloring often occurring in the peas of the 

 same pod. All were more or less purple spotted. These gave, in ' 

 Fo, 3 classes, the two grandparental types and the F^ type. The 

 statement regarding the presence of purple dotting on these Fg 

 segregates is rather obscure. 



Lock (53, p. 326) does not consider orange-brown testa color as 

 a separate character from gray-colored testa, and Bateson thinks 

 Corren's exceptional results in F^ of the cross just described may 

 be due to environment. The writer has distinctly orange-red seed 

 coat peas with white flowers in his collection from Chile and he 

 hardly beheves that present data justify Lock's contention, because 

 these peas do not mature as gray under the conditions in which ordi- 

 nary gray seed coat varieties have gray seeds. 



Colorless X dark brown seed coat varieties should according to 

 Tschermak's formula for at least one such variety [redfl. Kneifel- 

 erbse, S. 181 (86)] give all dark brown seed coat progeny in F^ 

 with or without purple dots, depending on the colorless variety used. 

 I have not, however, been able to find the published record of the 

 data upon which this formula is based. True breeding (86) dark 

 brown seed coat segregates crossed with colorless give dark brown 

 in Fj. 



Colorless X gray with maple pattern gives in F^ maple pattern 

 either with or without purple dots. The presence of the purple 

 dots in F^ of such a cross as this is altogether dependent on the kind 

 of colorless seed coat variety used, as the genetic evidence from 

 Bateson (i), Tschermak (86) and others shows that a gray maple 

 pattern seed coat variety may be crossed with colorless and give 

 maple and purple dots. The same maple variety may again be 

 crossed with a colorless, but this time a different one and give only 

 maple. Bateson (i) found British Queen to be a colorless seed 

 coat variety of the first type and Victoria one of the second type. 

 Tschermak [see Bateson (i)] secured 2 cases where Victoria X 

 unspotted varieties gave purple spots in F^, while reciprocals of the 

 same cross gave unspotted seed coats. In Tschermak's latest pub- 

 lication (86) on the subject, two varieties of Victoria are recognized, 



