WHITE— STUDIES OF INHERITANCE IN PISUM. 507 



conditions. Fruwirth (34), however, describes this pattern in the 

 Blauhiilsige variety as due to brownish, weak violet pigment 

 granules in the palisade cells. Lock says this pigment is easily sol- 

 uble in boiling water. 



Brown marbling or the maple pattern, as the English call it, is 

 associated only with colored flowers as far as the color is concerned. 

 Lock, however (54, 55), finds the pattern itself ("ghost mapHng") 

 without coloring, may be associated with white-flowered plants. The 

 brown pigment of the maple pattern is largely confined to the cell 

 walls of the outermost layer of I-shaped testa cells (55). The 

 pattern color deepens with age and is insoluble in boiling water. 



The "" eye " color pattern is characteristic of both colored and 

 white-flowered pea races. The color is present as a deep black at 

 the point of attachment, with a dark sooty tint usually present over 

 the seed as a whole. Some varieties have brown coloring (81) in 

 place of the black while other varieties are without color at this spot. 

 The brown hilum color according to Tschermak is always associated 

 with colored flowers and colored seed coats, so it may be consid- 

 ered as simply another of the numerous expressions of factor A. 



Violet eye is due to a violet hilum pigment, characteristic par- 

 ticularly of a race of Victoria peas with which Tschermak experi- 

 mented. 



Brown marbling, purple dotting or stippling and black eye may 

 all be associated in the same pea seed coat. In fact, a couple of 

 wild species obtained direct from Asia have seeds characterized by 

 all three of these color patterns. 



Varieties Studied. 



Varieties with colorless or almost colorless seed coats as de- 

 scribed under colorless : Griinbleibende Folger, Desirat, Auvergne, 

 Yellow-podded Sugar Pea, Express, Emerald, Victoria, Svalof Small 

 Green-seeded P'xsum, Prince of Wales (Tschermak, 81, 86) ; Griin 

 Spate Erfurter Folger (Correns, 14) ; Laxton's Alpha, Veitch's Per- 

 fection, Sunrise, British Queen, Victoria Marrow, Tres nain de 

 Bretagne, Earliest Blue, Ceylon Native No. i, Satisfaction, Ring- 

 leader (Lock, 54, 55, 56) ; Serpette, British Queen, Victoria Mar- 

 row, Ringleader, Nain de Bretagne (Bateson, et al., i), White-flow- 

 ered Mummy (Macoun, 57.5). 



