572 WHITE— STUDIES OF INHERITANCE IN PISUM. 



acter under favorable environmental conditions is completely hered- 

 itary. The variety was sent to Vilmorin from Switzerland in 1906 

 by M. Frommel and had emerald leaves. It has been suggested that 

 the absence of wax (glaucousness) has been partly responsible for 

 its origin, as the young growing peas in contact with each other, free 

 from wax, tend to grow together as do grafts. But in other emerald 

 varieties the peas do not adhere, so the attempted explanation is 

 not very satisfactory. 



Results from Crossing. 



Free seeds, glaucous foliage, pink flowers X chenille seeds, 

 emerald foliage, white flowers gave in F^, glaucous foliage, purple- 

 red flowers, and free seeds. In F„ a total of 175 progeny gave 144 

 with free seeds and 31 with adherent seeds or a ratio of approxi- 

 mately 4:1. Considering the combinations of this pair of charac- 

 ters with those of flower color and foliage character in F,, the 

 results were : 



-n.1 . 1 /ON r flowers colored 105 [ , . 



riants glaucous (1^0) -^ „ , . -{ seeds free 



" V o y ^ flowers white 33 [ 



f chenille 28 

 [flowers colored 29-^ . 



Plants emerald (39) \ ^ 



\ n , • of chenille 3 



I flowers white o J ^ 



[ free 5 



These results show all is in accordance with ordinary Mendelian 

 theoretical expectation both as to classes and numerical representa- 

 tion of classes, until the chenille and free seed pair of characters 

 is considered. Here one notes (i) that glaucous plants have only 

 free seeds whereas on a one-factor basis about 35 plants are ex- 

 pected to have chenille seeds; (2) that the chenille and free seed 

 characters are distributed among the emerald plants in approximately 

 3:1 proportions, but just the reverse of what ordinarily would be 

 expected, the dominant character in F^ in this cross being free 

 seeds. 



In F3 the Fo plants of various kinds tested out gave as follows: 



