646 FAMILY: TRICHOMONADID^ 



enter the sheath, in which situation they may be seen in incredible numbers, 

 either free or attached to the wall of the sheath by their fiagella. By 

 the end of the tenth day, before which the leech is not ready to feed 

 again, there are numerous flagellates in the proboscis sheath and anterior 

 part of the crop, but none farther back. Exceptionally, the fresh feed will 

 clear the leech of flagellates entirely, but, as a rule, the proboscis sheath, 

 which is emptied at the feed, becomes filled with the slender forms which 

 have resulted from multiplication of those left in the sheath or by a 

 further migration forwards from the crop. A fish, upon which infected 

 leeches had fed, first showed trypanoplasms in its blood on the seventh 

 day. According to Brumpt (19066), the trypanoplasms with which he 

 worked multiply in the gut, but do not invade the proboscis sheath. 

 Tanabe (1924) has given an account of the development of the trypano- 

 plasm of the Japanese loach {Misgurnus anguillicaudatiis) in the leech, 

 Hirudo nipponica. In the intestine active multiplication occurred during 

 the first three days, so that large numbers of small forms were produced. 

 After this there was a gradual disapjDearance of the flagellates, though in 

 some cases they persisted for eight or nine days. No mention is made of 

 any attempts to transmit the infection by means of these leeches. 



9. Family : Trichomonadid^. 

 The flagellates belonging to this family are characterized by the pos- 

 session of a variable number of flagella, a definite cytostome, and a rod- 

 like structure, the axostyle, which arises from the blepharoplasts and passes 

 through the body to its posterior end, through which it usually protrudes. 

 In some forms, one of the flagella is directed backwards, and its axoneme 

 may be attached to the border of an undulating membrane. In such cases 

 there is usually a stiff basal fibre, which lies along the line of attachment 

 of the undulating membrane to the body (Fig. 265). The family includes 

 the following genera: 



Genus: Trichomonas Donne, 1837. 



Ditrichomonas Cutler, 1919. 



Gigantomonas Dogiel, 1916. 

 ,, Eutrichomastix Kofoid and Swezy, 1915. 



Janickiella Duboscq and Grassi, 1923. 



Trichomitus Kofoid and Swezy, 1919. 



Devescovina Foa, 1905. 

 .» Foaina Janicki, 1915. 



Retortamonas Grassi, 1879. 



Protrichomonas Alexeiefl, 1911. 



Polymastix Biitschli, 1883. 



Hexamastix Alexeieff, 1912. 



Cochlosoma Kotlan, 1923. 



