ENDOMYCETALES 



141 



In other species which terminate the isogamous series, E. javanensis and 

 E. capsularis, the copulation branches have entirely disappeared (Fig. 21, 

 6-8). According to the conditions of environment, either the hyphal or the 

 sprouting condition may prevail. By swelling of the terminal cells of the 

 hyphae or by lateral sprouting (sometimes also intercalary), there arise 4- 

 spored asci. Each of these spores is divided into two unequal parts by annular 

 thickenings (Guilliermond 1909). Neither species is possessed of much fer- 

 mentative ability. 



From Endomycopsis two lines diverge. In Hansemda the ascospore is 

 cucullate, the mycelium has disappeared, although the cells are quite elongate 

 and sometimes in chains. Two of the sprout cells form copulation tubes to- 



Fig. 21. — Endomycopsis Lindneri. 1-5, E. capsularis ; 6-8, development of asci. (1, 2, 6-8 X470 ; 

 3-5 X500.) (After Mang-enot 1922 and Guilliermond 1909.) 



ward each other, the nuclei migrate into the bridge and fuse, the diploid 

 nucleus divides, both daughter nuclei migrate back into the cells, there divide 

 a second time, and develop two ascospores in each fusion cell. In Hansenio- 

 spora the cells of the sprouts mycelium are mostly citriform, sprouting only 

 from the poles. Copulation has not been observed and apparently the asco- 

 spores develop parthenogenetically. Pijper (1928) reports H. Guilliermondii 

 from the nails in a case of onychomycosis. 



In the other line diverging from Endomycopsis, the ring about the ascospore 

 occupies an equatorial position. In Williopsis saturnus we have vegetative 

 conditions very much as in Hansenula, where this species was formerly placed. 



