Art. II.] Herrick, Gustatory Paths in Fishes. 91 



The neurites from the cells of the lateral secondary gustato- 

 ry nucleus are in the main directed across the raphe through the 

 ventral commissure to reach the substantia reticularis grisea of 

 the opposite side of the oblongata (Figs. 31, 33). Here they 

 may come into relation with the dendrites of the nucleus am- 

 biguus and other motor nuclei. They are to be regarded as 

 direct reflex paths, differing from those provided by the inter- 

 mediary neurones in that there is no type II cell interpolated 

 between the root fiber and the tract neurone. Since this ar- 

 rangement prevails also for the main ascending gustatory path, 

 it is probable that this is the path of direct motor reflexes ex- 

 cited by the gustatory communis root fibers rather than by the 

 unspecialized visceral element. 



The dorsal and lateral groups of chief secondary neurones 

 receive by far the greater part of the vagal communis root 

 fibers, and since most of these root fibers are gustatory in func- 

 tion, we may infer with great confidence that the more lateral 

 group of secondary neurones is the chief path for direct gusta- 

 tory reflexes in the medulla oblongata and spinal cord, while 

 the more dorsal group is the main ascending or central gusta- 

 tory path. 



Neurites from the dorsal group of secondary neurones pass 

 through the middle of the vagal lobe in slender compact bun- 

 dles, directed downward and outward to a position ventrally of 

 the spinal V tract. Here they turn and take a longitudinal 

 direction, most turning cephalad and constituting the ascending 

 secondary gustatory tract ("sekundare Vagusbahn," Mayser), 

 but some caudad. The latter are derived chiefly from the more 

 lateral part of the vagal lobe and more probably should be rele- 

 gated to the cells of the more lateral group of neurones. The 

 relations of the ascending fibers from the vagal lobe and both 

 the ascending and the descending fibers from the facial lobe are 

 indicated in Fig. 37. 



The relations of the cells of the intermediary nucleus of 

 the vagal lobe are not very clearly brought out in my prepara- 

 tions. Figs. 29 and 34 show their dendrites in relation with 

 the neurites of type II cells of the vagal lobe. Some prepara- 



