Proceedings of 

 the United States 

 National Museum 



SMITHSONIAN INSTITUTION • WASHINGTON, B.C. 



Volnme 111 I960 Number 3425 



NOTES ON LARVAE OF NINE GENERA OF APHODIINAE IN 

 THE UNITED STATES (COLEOPTERA : SCARABAEIDAE) 



By Manohar Lal Jerath^ 



The coprophagus scarabaeid subfamily Aphodiinae contains a great 

 many species of small to medium-sized beetles with a great diversity 

 of habits. Most species are found in dung; some, however, are found 

 in soil or sand feeding on organic matter or roots of living plants; 

 others are said to be parasitic. The subfamily is worldwide in 

 distribution. 



This study of the systematics of larval Aphodiinae, begun in 

 November 1954, was undertaken because practically nothing was 

 known of the American genera and species. It was suggested by 

 Dr. Paul O. Ritcher, Department of Entomology, Oregon State 

 College, Corvallis, Oreg., and is based on the study of larvae and 

 adults loaned from the U.S. National Museum (USNM), Dr. Ritcher's 

 personal collection (POR) and material collected by the writer. The 

 assistance and encouragement of Mr. O. L. Cartwright of the U.S. 

 National Museum are gratefully acknowledged. 



' Hoshiarpur, Punjab, India. 



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