236 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 



/porter! Group/ 



A 



/pgllidipennis Groupy 

 colei 



eugonotus Group/ /borealis Group/ 



Figure 2. — Phylogenetic tree of Ogcodes species groups. 



features of both the pallidipennis and colei groups, accounts for my 

 placing it as ancestral to the two last-named groups. The species of 

 the pallidipennis group have all retained vein Mi but have lost 

 crossvein m-cu, and have developed stronger, well-built male genitaUa. 

 Members of the colei group have retamed (or lost) one or both of the 

 wing veins, and have, for the most part, much-reduced male genitalia. 

 From the colei group in the Nearctic region arose the monotypic 

 subgenus Neogeodes, judging from its more reduced wing venation, 

 male genitalia, loss of the antennal style, and the subsequent reduction 

 in size of the terminal antennal segment. The porteri group has greatly 

 reduced wing venation, but because the monotj^pic species is known 

 from only one specimen, which did not possess antennae and whose 

 male genitaha could not be examined, its placement and rank are 

 both questionable. However, judging from the known distribution 

 and wing venation, it probably represents a highly evolved group which 

 was derived from a Chilean species of the eugonatus group. 



In the phylogenetic tree for the Nearctic species (text fig. 3), only 



