NO. 1976. • TREE SHREW 8: FAMILY TUPAIID^—LYON. 45 



TUPAIA GLI3 GLIS (Diard). 



1820. Sorex glis Diard, Asiat. Jonm. Month. Reg., vol. 10, p. 478, November, 

 1820. 



1821. Tupaiafcrruginea Raffles, Trans. Linn. Soc. London, voL 13, 1822, p. 256, 

 published May, 1821. 



1822. Sorex glis, Diard and Duvaucel, Asiat. Res., vol. 14, p. 472, pL 9. 

 1911. Twpaia ferruginea penangensis Robinson and Kloss, Journ. Fed. Malay 



States Mus., vol. 4, p. 242, November, 1911. (Type No. 1445/11, Selangor 

 Museum, collected at Telok Behang, Penang Island, Apr. 2, 1911.) 



Type-locality. — Penang Island, off west coast of Malay Peninsula. 



Tliere is no type-specimen of Twpaia glis glis. Penang is the only 

 exact locality mentioned in the original description, and consequently 

 is regarded here as the type-locahty. No mention of its occurring 

 on the Malay Peninsula is made. Of course Diard and RafHes had 

 the same animal in mind in describing their respective species, but 

 the practical results are that glis is the name for the Penang animal 

 Mid ferruginea for the Sumatran and peninsular one. 



Geographic distribution. — Penang Island. See No. 21 on map on 

 page 75. 



Diagnosis. — A sHghtly differentiated geographic form of Tupaia 

 glis, differing in smaller size and a duller paler coloration of the upper 

 parts. 



Color. — The general stylo of coloration of Tupaia glis glis is very 

 similar to that of wide ranging T. glis ferruginea, hut the upper parts 

 of body instead of being a fine grizzle of ferruginous or chestnut and 

 blackish, is a grizzle of rather dark tawny or tawny ochraceous and 

 blackish, while the head and neck are a grizzle of ochraceous buff 

 and blackish. The tail is generally duller in T. glis glis. In other 

 respects tlie two animals are essentially the same. 



STcull and teeth. — -The skull and teeth of Tupaia glis glis are dis- 

 tinctly smaller than those of T. glis ferruginea, especially seen in the 

 brain case. (Plate 8, fig. 8.) 



Measurements. — The usual measurements of adults: Head and 

 body, 1G5 mm.; tail, 155; hindfoot, 42-43; condylo-basal length, 47; 

 Z3^gomatic width, 25; width of braincase, 19; maxiUary tooth row, 

 18.5; most of them agreeing with the minimum measurements of 

 Tupaia glis ferruginea. For measurements of all specimens examined 

 see table, page 46. 



Remarlcs. — Tupaia glis glis is closely related to T. glis ferruginea 

 of the neighboring mainland of which it is zoologically a geographic 

 form. Robinson and Kloss were entirely right in describing it as a 

 subspecies of the mamland animal, and it is perhaps a misfortune that 

 glis was not originally used for the mainland form, or rather that 

 ferruginea was not proposed first, as the latter name has been so long 

 in use that it seems a pity to have it rank as a subspecies. 



Specimens examined. — Seven in the British Museum, and two loaned 

 by the Selangor Museum. See table, page 46. 



