NO. 1976. 



TREESHREW8: FAMILY TU P All D^— LYON. 



121 



§ 



stuffing the head, show ill-defined but still evident light Imes over and 

 below the eye, thus suggesting the more pronounced face markings 

 seen in most species of Dendrogale. A shoulder stripe is present. 

 Mammae are 3-3 = 6, 1 axillary, 1 inguinal, 1 ventral near the ingui- 

 nal one. (Plate 4.) 



Cranial characters. — The skull of Anathana shows little devia- 

 tion from the typical Tupaia skull. The rostrum is short and heavy, 

 does not arise from the skull abruptly as in the Twpaia minor group, 

 where the rostrum is also relatively short. The distance from the 

 lachrymal notch to end of premaxilla is about equal to the distance 

 from that notch to end of external pterygoid plate. In Tupaia, with 

 the exception of the minor group, the dis- 

 tance from the lachrymal notch to end of 

 premaxilla equals the distance from the 

 notch to the external auditory meatus. 

 Correlated with the shortened rostrum 

 the premaxilla is correspondingly short, 

 antero-posteriorly and relatively high, 

 supero-inferiorly. Usually in Tupaia the 

 naso-premaxillary suture is considerably 

 lengthened or apparently obliquely pushed 

 backward, making the premaxUla a rather 

 obliquely elongated bone. In Anathana 

 the premaxilla is somewhat quadrate in 

 shape and only slightly distorted. Ex- 

 treme obliquity of the premaxilla is seen 

 in the genus Tana. The fenestra in the 

 zygoma in AnatJiana is reduced to a small 

 oval foramen quite different from the 

 large opening found in Tupaia. While 

 it appears a trivial character, yet m the 

 Tupaiidse the size of this openmg is an 

 important feature correlated with other pecuKarities 

 page 120. 



Dental characters. — The most evident of the dental characters in 

 Anathana is the relatively small size of the lower canine, so that when 

 the lower jaw is viewed from the side this tooth does not stand con- 

 spicuously higher than the adjacent incisor and premolar. The devel- 

 opment of hypocones on the first and second upper molars is very con- 

 spicuous in Anathana; they are larger than in any other member of 

 the family though nearly equalled by the hypocones on the first two 

 upper molars of Urogale and Ptilocercus. All the upper molariform 

 teeth of Anathana are relatively wider and shorter than they are in 

 the other genera. This is especially well shown m the third and 

 fourth premolars, where the protocones are better developed and 

 extend inward toward the median line more than is the case in other 

 genera of Tupaiidse. Looked at on its grindmg surface the fourth 



Fig. 7.— Upper and lower tooth- 

 rows OF Anathana wkoughtoni 

 Type x 3, Reg. No. 96. 11. 7. 1, 

 British Museum, Mandvi, India. 



See fig. 6 on 



