207 



reasons, an intimate alliance, differing chiefly in their qualities, in the non- 

 connivence of their anthers, the aestivation of the corolla, and the structure 

 of the ovarium. To Lorantheae they also approach, but differ in the relation 

 of the anthers to the lobes of the corolla, and in other points. But if we 

 consider the tribe called Sambucinese, our view of the affinities of the order 

 will take a different turn, and we shall find an approach to an order the 

 relationship of which would hardly have been suspected, viz. Saxifrageae : 

 this is established through the intervention of Hydrangea, a genus usually 

 referred to Saxifragei3e, but which it appears more advisable to station by the 

 side of Viburnum, from which it is undistinguishable in habit, and with which 

 it accords in inflorescence and in the constant disposition of its flowers to 

 become radiant, but which differs in being polypetalous and polyspermous. 

 Besides these points of affinity, Caprifoliacese probably tend towards Umbel- 

 liferae through Sambucinese. 



The following are the characters of the sections, if they be sections, of 

 this order : — 



1. Lonicere;e. The Honeysuckle Tribe. 



I/jnicerese, Ach. Rich. EUm. de la Bot. ed. 4. 484. (1828). — Caprifoliea^, 



Dec. and Duby, 244. (1828.) 



Corolla tubular. Berry 2- to 4-celled, with 1 or many-seeded cells. 

 Style 1. Leaves opposite. 



True Caprifoliacese are said by Mr. Brown to be distinguished from the 

 other genera hitherto associated with them, in the raphe being on the outer 

 instead of inner side of the ovulum. Brown in Wallick, PL As. p. 15. 



Examples. • Caprifolium, Lonicera, Linnaea, Abelia, Triosteum, Dier- 

 f ilia, Schopfia. 



2. Sambucine*. The Elder Tribe. 



Sambucineaj, A. Rich. Diet. Class. 3. 173. (1823); Dec. and Duby, 244. 



(1828); Link Handb. 1. 662. (1829.) 



Corolla rotate. Ovarium 3- or 4-celled, with solitary pendulous ovules. 

 Styles 3 or 4. Flowers in cymes, the lateral ones often radiant. Leaves 

 opposite. 



These pass into Lonicereae through Viburnum davuricum, which has 

 the tubular corolla of a Lonicera, and into Hydrangeaceae through the 

 radiant-flowered species of Viburnum. With Hedera they are connected 

 through Cornus. 



Examples. Viburnum, Sambucus. 



3. Hederace.e. The Ivy Tribe. 

 Hederaceae, Ach. Rich. Bot. Med. 2. 449. (1823); Dec. and Duby,' 244. 



(1828.) 



Corolla polypetalous. Disk epigynous. Style 1. Drupe or berry with 

 1-seeded cells. Leaves opposite or alternate. 



Ach. Richard considers this a distinct order, on account of its polypetalous 

 corolla and epigynous disk. 



Examples. Hedera, Cornus. 



4. Hydrangeace^. The Hydrangea Tribe. 



Corolla polypetalous. Styles 2 to 5. Fruit succulent or capsular, 2- to 

 3-celled, many-seeded. Leaves opposite. Flowers in cymes, the lateral ones 

 often radiant. 



The characters of this tribe are so strongly marked as to justify its being 

 established as an independent order ; but the habit of the species is so entirely 

 that of Viburnum, that I am not willing to separate them without absolute 

 necessity. There is a remarkable resemblance between their seeds and those 

 of Begonia. 



Examples. Hydrangea, Adamia» . . 



