COMl'OSIXlOX OF THli FL()1!A. 



85 



recent Aiistraliiin spcvies, is re])resoiitecl by 11 

 fossil species iu tiie ()li<i;oceiu' of Kuro])c, in 

 France, the T_vto1, Saxony, and Cire(n'(>, and hy 

 no less than 17 species in the Mit)cene of France, 

 Italy, Switzerland, Baden, llesse, I'rnssia, Bo- 

 hemia, Austria, Styria, Croatia, and Hungary. 



The geiuis Knightia 11. Brown, which in- 

 cludes a nioderii species iu Australia and 2 in 

 New Caledonia, includes a fossil form in the 

 Oligoceuc of Australia and anotlun' in Graham 

 Laud (Antarctic Continent)' in beds which are 

 regarded as Oligoceuc. The allied geiuis 

 Kniglitites Saporta contains 2 species from lh(> 

 Sannoisian of France. 



The remai-kable genus Lomatia, previously 

 mentioned, is represented by 4 existing s))e- 

 cics in Australia, 2 in Tasmania, and .3 in Chile. 

 As might be expected from these moilern iso- 

 lated occurrences, there are over 30 fossil spe- 

 cies, some of them based on associated leaves 

 and fruits. The ohh^st of these species are 2 

 (perhaps wrongly identified) in the Dakota 

 sandstone. Eocene forms iuchule species from 

 the Green River formation of North America, 

 an Ypresian species from the south of England, 

 and an Italian species. There are about IS 

 Oligoceuc species, some of which are very 

 characteristic. They occur in t!ie Tyrol, 

 Saxony, Baltic Prussia, Styria, Australia, and 

 Tasmania, and the relatively large number of 

 4 are recorded by Dusen from Graham Land. 

 The Australian and Tasmanian forms may he 

 Miocene instead of Oligocene. About a dozen 

 Miocene species have also been recorded from 

 such widely separated areas as Colorado, Swit- 

 zerland, and Carniola. The characteristic de- 

 tails seen in the wonderfully preserved leaves 

 in the volcanic ash beds at Florissant, Colo., 

 from which 7 forms of Lomatia have been de- 

 scribed, leave no doubt as to the validity of 

 the generic identification. 



The allied genus Lomatites Saporta is rep- 

 resented by a Cenomanian species in Saxony 

 and 5 or Oligocene species in France. The 

 genus Stcnocarpus R. Brown, of which 1 1 ex- 

 isting species have been found in New Cale- 

 donia and 3 additional species at other places, 

 ranging from North Australia to New South 

 Wales, has a siiigle fossil species in the Oligo- 

 cene of Saxouv. 



I Du.sdn, P. C. U., Wissenschaftliche Ergebnisse der Schwedischen 

 Sudpolar-Expedilion, 1901-1903, Bd. 3, Lief. 3, p. 7, pi. 1, figs. 7, 9, 11, 

 190S. 



The genus Persoonia Smith contains 60 ex- 

 isthig s])(M-ies iu Australia and 1 in New Zea- 

 land. '!'hc fossil record includes 2 widely dis- 

 tributed s])ccies in the L"])pcr Cretaceous of 

 North America; 1 in the Eocene of England; 

 4 in llu> Oligocene of Tyrol, Saxony, Styria, 

 and Greece, and 10 in the Miocene of France, 

 Italy, Swifzerland, Baden, Bohemia, Styria, 

 Croatia, Carniola, and Slavo)iia. Deane records 

 a Tertiary species from New South WsUes. A 

 large number of these fossil forms of Pi-rsoonia 

 are not especially convincing, bn( cei-tainly the 

 tlirei> Euro])ean species /-'( rsixiniu ciispidatd, P. 

 dnplines, and P. iin/rfilliis of Ettiugshausen,- 

 which have the leaves associated with char- 

 acteristic fruits, are above suspicion. 



Bowerbank, iu his classic study of the pyrit- 

 i/.ed fruits and seeds from the Isle of Sheppey, 

 established a genus which he called Petro- 

 philoides from its resemblance to the genus 

 Petrophila R. Brouni, which inchules about 35 

 existing species in Australia, most of which 

 are confined to West Australia. Bowerbank 

 described several species, one of which was 

 showm by .Starkie Gardner to be an ^Unus fruit, 

 and others have been referred to Se(iuoia. 

 Ettmgshausen^ in the study of the Sheppey 

 fruits, after careful comparisons, retained three 

 English Eocene species and the genus has also 

 been recognized in the Sannoisian of Ualmatia 

 and Styi'ia. 



The genus Leucadenclrites was established 

 by Saporta for a Sannoisian species of south- 

 eastern France from its resemblance to Leuca- 

 deudron Ilennann, which includes moi-e than 

 70 existing species in South Africa. 



The genus Grevdlea R. Brown contains 56 

 existing species which are confined to Australia. 

 The fossil record includes a Cretaceous species 

 in Australia; 2 Cenomanian species in Bohemia 

 (Gre\'illeophyllum Velenovsky); 3 Eocene spe- 

 cies in England, France, and Italy; 12 Oligo- 

 cene species, mostly in southern France but 

 also represented in Saxony, Tyrol, Bohemia, 

 Styria, and Greece; and 12 Miocene species in 

 France, Switzerland, Bohemia, and Croatia. 



The genus Embothriuni Forster, already 

 alluded to, includes 4 existing species in South 

 America, wliich range from Chile to thci Straits 

 of Magellan, and 1 species in Australia. This 



2 Ettingshausen, C. von, K. .\kad. Wiss. Wien, Math.-Nat. CI., Sitz- 

 ungsbcr., Bd. 7, pp. 71S-719, pi. 30, figs. 5-11, 1831. 

 ' Koy. Soc. London Proc, vol. 29, p. 394, 1879. 



