MYRTALES. 



319 



Occurrence. — Lagrange formalion (in l)C(ls of 

 Wilcox age), Piirvpar, Ilonrv Couiify, Tcnn. 

 (collected l)y E. W. Bern). 



Collection. — U. S. National Muscinn. 



Eugenia grenadensis Benv, n. sp. 

 Plate XCI, figures 4 and "i. 



Description. — Leaves medium sized for tiiis 

 genus, lanceolate in general outline, wid(>st at or 

 slightly below the middle, fiom wliieli ])oint 

 they taper ahnost equall\- in hotii directions in 

 the smaller leaves, although iu the larger leaves 

 the base is wider than the tip. Apex abru])lly 

 pointed. Base euneate. Lengtli ranges from 

 5.5 to 7 centimetere. Maxmium width, at or 

 slightl}' below the middle, ranges from 1.5 to 

 2 centimeters. Margins entire, invariably 

 somewhat irregularly undulate in the material 

 collected. Texture coriaceous. Petiole short, 

 very stout, and much curved, al)out 3 milli- 

 meters in length. Midrib stout throucrliout its 

 length, promiiuMit. Secondaries numerous, 

 thin, brochidotlrome ; thej- diverge from the 

 midrib at acute angles of about 35° and curve 

 but slightly m their ascending course, bemg 

 scarcely distinguishable from the tertiaries and 

 not at all difl'erentiated in the upper part of 

 the leaf. Tertiaries thhi, forming a close-set 

 ascendmg transverse areolation. 



This species is well distinguished among the 

 Wilcox species of Eugenia, bemg longer and 

 narrower than Eugenia liilgardiana Berry, which 

 also differs in its thicker midrib, fewer, stouter 

 secondaries, and tertiary areolation. The other 

 Wilcox species, Eugenia densinervia (Lesquereux) 

 Berry, is a small linear-lanceolate falcate form 

 quite unlike Eugenia grenadensis. The leaves of 

 a number of existmg species of Eugenia closely 

 resemble those of the fossil species. Those that 

 reach our southern coast are as a rule relatively 

 wider, but Eugenia confusa De Candolle is not 

 very dissimilar from the fossil form. 



Occurrence. — Grenada foi'mation, Grenaila 

 Grenada County, Miss, (collected by E. N. Lowe 

 and E. W. Beri-y). 



Collection. — U. S. National Museum. 



Genus CALYPTRANTHES Swartz. 

 Calyptkantiies eocenica Berry, n. sp. 



Plate XC, figure 5. 

 Description. — Rigid coriaceous leaves, ellip- 

 tical or ovate in outline, the base broadly eune- 

 ate or rounded, and the tip acute and not ex- 

 tended. Length about 7 centimeters. Ma-xi- 



nuun widtii, in the middle^ ])art of the leaf, 

 al)out 3.5 centimeters. Midrii) rather stout. 

 Secondaries tliin, numerous, close set, subpar- 

 allel, in |>laces forked and anastomosing; tliey 

 (hvei-ge from tiie midrib at wide angles and ])ui'- 

 sue a neai'Iy straight course almost to tiie mar- 

 gin, wliere their ends arc joined by a marginal 

 vein parallel willi \\\e entire niai-gin. 



Tiiis characteristic new species res(>nd)les a 

 number of existing American gen(>ra of the Myr- 

 tacea^ as well as several West Indian species of 

 .Sapotacea^, especially of tlie genus Ciirysopiiyl- 

 lum. It also resembles several species of the 

 myrtaceous genus Aulomyrcia Berg (Myrcia 

 De CandoUe) from tropical South America, but 

 is especially close to certain species of Calyp- 

 tranthes, particulai'ly Cdhjptraiithes sgzi/gium 

 (Linne) Swartz, which is a sJn-ub or small tree 

 20 or 30 feet in height ranging from sea level to 

 the mountains in rich woods on the West In- 

 dian islands, and mostly an upland form. 

 Some modern students refer it to the allied 

 genus Chytraculia R. Brown. The fossil is 

 near the maximum in size of the leaves of this 

 species and except for its more sharply pointed 

 tip is identical with the leaves of this recent 

 species m all its characters. 



Among previously described fossil forms it is 

 perhaps most like the somewhat smaller Myrtus 

 rectinenns, which is described by Saporta' 

 from the Oligocene (Tongrian) of St. Zacharie 

 in southeastern France. 



The genus Calyptranthes appears to be an 

 early branch of the Eugenia-Myrcia plexus, 

 supposed to have been largely developed in the 

 American tropics during the Cretaceous. As 

 treated by Niedenzu- it comprises about 70 

 species, rangmg from Mexico and the West In- 

 dies to southern Brazil. Several insular forms 

 have been referred to this genus on evidence 

 which I b(>lieve is insufficient. These arc 3 forms 

 from the Fiji Islands, 1 from Africa and Mauri- 

 tius, and 1 from Java. With these exceptions 

 the genus is entu-ely American in the existing 

 flora. An undescrilied fossil species is present 

 in the Oligocene of the Isthmus of Panama. 



Occurrence. — Lagrange formation (in beds of 

 Wilcox age), Puryear, Henry County, Tenn. 

 (collected by E. w". Berry) . 



Collection. — L^. S. National Museum. 



' Saporta, G . do, Etudes siir la vc-'gi^tat ion du siid-cst do la France h 

 l't'i>0(|uc lertiaire, vol. 1, p. 251, pi. 11, fig. 5, l.^il3. 

 • Englcr, A., and I'rantl, K., Die natiirlichen I'llaiizeiifamilien, 1S93. 



