HEATH FAMILY 47 



(1903), type loe. Gasquet (w. of), Del Norte Co., Howell. A. subcordata Eastw., Lflts. W. Bot. 

 1:61 (1933), type loc. Santa Cruz Isl. (w. end), J. T. Howell 6335; leaves subeordate at base or 

 acute. A. cmstacea Eastw., I.e. 1 :74 (1933), type loc. Kings Mt., San Mateo Co., Scale. A. rosei 

 Eastw., I.e. 1 ; 77 (1933), type loc. Lake Merced, San Francisco peninsula, L. S. Rose. A. zacaensis 

 Eastw., I.e. 1:79 (1933), type loc. Zaea Lake, Santa Barbara Co., Eastwood 681. A. cushingiana 

 Eastw., Lflts. W. Bot. 1:75 (1933), type loc. Mt. Tamalpais (south slope), "abundant on lower 

 slopes," Eastwood 11,075a; not at all or only slightly glandular; branchlets canescent, not bristly 

 or only slightly so. [A. glandulosa var. crassifolia Jepson, Man. 749 (1925), lapsus calamitosus 

 typographicus.] 



19. A. tomentosa Lindl. Explorers Manzanita. Shrub 3 to 4 feet high, the 

 erect stems with shreddy bark, several or many from a broad woody root-crown ; 

 branchlets pubescent or tomentulose, more or less glandular, very leafy ; leaf -blades 

 oblong-ovate, mostly acute, mostly truneatish at base, green and glabrous above, 

 a fine tomentum or close felt-like pubescence beneath, % to 1 (or 2) inches long; 

 petioles short ; panicle compact (^ to V2 or 1 inch long) , often reduced to a raceme ; 

 berry depressed-globose, glabrate or with scattered white liairs, 4 lines broad ; nut- 

 lets distinct or some united, rugulose and somewhat ridged. 



Sandy ridges or hills, 10 to 1000 feet : Santa Cruz Mts. (w. slope) ; northern 

 Monterey coast line region ; Sau Luis Obispo coast ; Santa Cruz Isl. ; south to the 

 San Diego coast line in a varietal form. Jan.-Mar. 



Tax. note.^ — Arctostaphylos tomentosa is common in the neighborhood of Monterey. The 

 shrubs possess thick woody root-crowns from which sprouts arise after fire or mutilation. This 

 species, first described as Arbutus tomentosa by Frederick Pursh, rests upon specimens gathered 

 by Menzies on the "Northwest Coast." Menzies' specimens are in the Herbarium of the Natural 

 History Museum, South Kensington, London. The particular sheet used by Pursh bears seven 

 small branchlets of unequal size and represents a mixture of two forms, first, a form with tomen- 

 tulose leaves, second, a form with stiffish-hirsute branchlets. The first form consists of two 

 branchlets mounted diagonally across the sheet; they have the leaves finely tomentose beneath. 

 The second form consists of five branchlets, the branchlets bearing long stifSsh spreading hairs 

 which arise from a dense tomentulum, the leaves acute or acutish at apex and base (at most ob- 

 tuse or very obtuse at base), not tomentulose beneath but only puberulent, and subglabrate above. 

 The two branchlets first spoken of correspond better on the whole, perhaps, than the other five 

 branchlets, with Pursh's original description, which itself must have covered in part at least the 

 two lots of specimens. The tomentose form matches in leaf shape and size and in tomentum very 

 exactly Arctostaphylos vestita Eastw. of Monterey, a fact which has been previously pointed out 

 by Alice Eastwood and C. V. Piper. The branchlets are quite densely white-tomentulose and there 

 are no hirsute or long spreading hairs. The more well developed leaf -blades have a subeordate 

 base (Jepson, Types and Original Citations, 1:68-71. 1926. nis.). Since Arctostaphylos vestita, 

 in just the form in which it occurs at Monterey, does not exist elsewhere on the coast line of Pacific 

 North America so far as known, it would seem likely that Menzies on one of his visits to Monterey 

 between 1791 and 1795 collected his specimens of A. tomentosa at that point, not only the tomen- 

 tose branchlets referred to above, but also the remaining five branchlets of the type sheet which 

 are here regarded as variants of A. tomentosa, variants readily duplicated by latter day collections 

 at Monterey. In that early day, moreover, the term "Northwest Coast" was used loosely and some- 

 times included northern California. It is of course possible that the specimens, although collected 

 at Monterey, may have been mislabeled. Indeed many of Menzies' specimens were not labeled 

 at all as to precise locality and are known only as belonging to the general collection made on the 

 "Northwest Coast." According to the Northwest Coast portion of Menzies' Journal of Vancouver's 

 Voyage, AprU to October, 1792, edited by C. P. Neweombe (Archives of British Columbia, Mem. 

 5:20, — 1923), Menzies discovered a species of Arctostaphylos at Port Discovery on the present 

 Washington coast. In so far as the evidence is now understood, it may be said that these Port 

 Discovery specimens were not the ones used by Pursh. It may also be added that recent collec- 

 tions made at Port Discovery Bay, Clallam Co., Wash. {Geo. Neville Jones) show that the shrub 

 of that locality (now referred to Arctostaphylos Columbiana Piper) is quite different from the 

 shrub at Monterey which was published as Arctostaphylos vestita Eastw. 



Locs. — Santa Cruz Mts.: n. of Santa Cruz, Jepson 9779. Monterey Co.: Del Monte Heights, 

 Jepson 5700; Monterey, Jepson 4004; Huckleberry Hill, Cooper 198. San Luis Obispo Co.: San 

 Simeon, K. Brandegee. Santa Cruz Isl.: Pelican Bay (ridge above), Jepson 12,083 (branchlets 

 weakly bristly). 



Variation note. — Arctostaphylos tomentosa is an aggregate species exhibiting a swarm of 

 varieties or forms chiefly marked by differences in hue of foliage and presence or absence on leaves 

 or branchlets of two different kinds of trichomes, short soft hairs and bristly hairs, in various 

 combinations and either non-glandular or glandular. Some of these forms which have received 

 n.ames are here noted : Shrubs having branchlets with bristly spreading often glandular hairs 



