GILIA FAMILY 143 



8292; betw. Silver Canon and Big Prospector Mdw., White Mts., Jepson 7246; Mono Mills (near 

 Mono Lake), Jos. 4" Hilda Grinnell 1060a; Grass Valley, Lassen Co., Fischer 4" Jolaison 1202; 

 Egg Lake, Modoc Co., M. S. Baker. 



Var. hallii (Parish) Jepson. Low erect shrub 10 to 20 inches high, the stems somewhat vir- 

 gate; branchlets on the 2 or 3-year-old stems short, bud-like, 4-angled and thus lycopodioid in 

 aspect; leaves usually opposite, palmately 3-parted, the middle lobe 2 to 7 times longer than the 

 lateral ones ; corolla white or yello-nish, 6 to 10 lines long, the lobes narrowly obovate. — Mountains 

 on northwest and west sides of the Colorado Desert, 2000 to 6000 feet: Pinon "Well Canon, north 

 of Indio, Jepson 6003; Coyote Caiion, Santa Rosa Mts., Jepson 1456; Mountain Sprs. (summit 

 above), e. San Diego Co., Jepson 11,805. Apr.-May. 



Var. shastense Jepson var. n. Erect shrub, 13 to 16 inches high ; branches and branchlets 

 very erect, the somewhat remote leaves all opposite quite to or nearly to the top ; corolla white. — 

 (Fruticulus erectus, strictus; folia subremota, plerumque opposita; corolla alba.) — Mayten (e. 

 of). Little Shasta Valley, Siskiyou Co., Jepson 19,994 (type). 



Var. subflavidum Jepson var. n. Shrub 1 to 2 feet high and 2 to 3 feet broad, the shoots of 

 the season puberulent, the leaves alternate, rather remote ; leaf -lobes strongly unequal ; corolla 

 pale yellow. — (Frutex 1-2 ped. altus, 2-3 ped. latus, caulibus in praesenti anno puberulenti- 

 bus; folia alterna subremota; foliorum lobi inaequales; corolla subflavida.) — Rocky canon walls, 

 4500 to 5500 feet: desert ranges from the eastern Mohave Desert (Fountain Caiion, Providence 

 Mts., Mary Bcal 665; Bonanza King Canon, Providence Mts., Jepson 18,280), north to Inyo Co. 

 (Leadfield, Grapevine Mts., Jepson 19,830, type) and Mono Co. (Long Valley, Almeda Nordyhe). 



Refs. — Leptodacttlon pungens Jepson, Man. 806, fig. 779 (1925). Cantua pungens Torr., 

 Ann. Lye. N. Y. 2 :221 (1826), type loc. "Valley of the Loup Fork?," Neb., James. Gilia pungens 

 Benth.; DC, Prodr. 9:316 (1845). Phlox hookeri Doug].; Hook., Fl. Bor. Am. 2:73, pi. 159 (1838), 

 type loc. near the narrows of the Oakangan and Priest's Rapid of the Columbia River, Douglas. 

 Cantua. pungens var. hookeri Howell, Fl. Nw. Am. 453 (1903). L. pungens var. hookeri Jepson, 

 Man. 807 (1925). Gilia pungens subsp. eupungens var. hookeri Brand; Engler, Pflzr. 4-^": 126 

 (1907). Gilia pungens var. squarrosa Gray, Proc. Am. Acad. 8:268 (1870), "arid districts of 

 Nevada and Utah," Anderson, Watson. CantiM pungens var. squarrosa Howell, Fl. Nw. Am. 453 

 (1903). L. patens Hel., Muhl. 1:146 (1906), based on Gilia pungens var. squarrosa Gray. Gilia 

 tenuiloba Parish, Erythea 7:95 (1899), type loc. San Jacinto Mts., Parish 689 ("probably Tah- 

 quitz Valley"). G. pungens var. tenuiloba Mlkn., Univ. Cal. Publ. Bot. 2:43 (1904). L. tenuilo- 

 bum Hel., Muhl. 1:146 (1906). L. pungens var. tenuilobum Jepson, Man. 807 (1925). Gilia 

 pungens subsp. pulchriflora Brand, Ann. Conserv. et Jard. Bot. Genfeve, 15-16:333 (1913), type 

 loc.Farewell Gap, Tulare Co., Culbertson. G. jaegeri Munz, Bull. S. Cal. Acad. 31:68 (1932), 

 type loc. Tahquitz Peak, San Jacinto Mts., Jaeger. Var. hallii Jepson, Man. 807 (1925). Gilia 

 hallii Parish, Erythea, 7:94 (1899), type loc. Coyote Canon, sw. slope of El Toro Mt., Santa Rosa 

 Mts., Hall. G. pungens var. hallii Mlkn., Univ. Cal. Publ. Bot. 2 :42 (1904). L. hallii Hel., Muhl. 

 1:146 (1906). Gilia pungens subsp. hallii Brand; Engler, Pflzr. 4"0:128 (1907). Var. shas- 

 tense Jepson. Var. subelavidum Jepson. 



2. L. califomicum H. & A. Prickly Gilu. Erect shrub (or at least woody 

 below), 1 to 5 feet high, the branches thickly clothed with spiny foliage; stems and 

 calyces puberulent; leaves usually glabrous or glabrate, mostly deciduous, alter- 

 nate or occasionally opposite, especially below, 1 to 3i/4 lines long, palmately 5 to 

 9-divided into rigid narrowly linear acerose lobes ; flowers sessile, solitary in the 

 closely approximate upper axils, thus congested into few-flowered clusters; calyx 

 % to as long as corolla-tube, its subulate lobes pungent, equal or nearly equal, % 

 to % as long as calyx-tube, the hyaline intervals broad ; corolla salverf orm, bright 

 rose-color or pink, 7 to 9 lines long, 9 to 17 lines wide, its broadly obovate lobes 

 equaling or slightly exceeding the tube; stamens included, inserted above the middle 

 of the tube, the filaments very short; capsule elongated, the cells many-seeded; 

 style rather short, included. 



Chaparral slopes or borders of washes, 600 to 5000 feet : in the outer and middle 

 ranges of San Luis Obispo Co. ; coastal Southern California from Santa Barbara 

 Co. to Orange Co. Peb.-May. 



Field note. — From the standpoint of its ecological interest and its showiness in flower, Lepto- 

 dactylon califomicum is one of the more remarkable plants of the Californian flora. Commonly 

 a member of the chaparral or chamise formations on the driest or most sterile slopes, its salver- 

 form corollas and prickly foliage at once attract the attention of the traveler. In the canons of 

 the Purisima Hills, for example, the shrubs, 3 to 5 feet high, are covered in April vrith large clus- 

 ters of satiny flowers of a rose pink or Godetia hue which glorify the open slopes and form an 

 enlivening contrast to the dull tone of the chamise formation. 



