GILIA FAMILY 145 



the calyx-lobes. In a single individual of Navarretia cotulaefolia calyces have been observed to 

 vary as follows: (1) lobes or segments 4 or 5, short, subequal, entire; (2) lobes similar to 1, but 

 more unequal; (3) lobes unequal, 1 long and toothed, 4 shorter and entire; (4) similar to 3, but 

 two of the lobes toothed, one of these often with approximate and divergent teeth at base; 

 (5) usually 3 of the lobes toothed, one of these much toothed and sometimes bipinnatifid. This 

 last form, a bipinnatifid segment, is similar to the smaller and shorter bractlets which subtend the 

 flower ; and these inner bractlets pass by every gradation in size and decrease of segmentation into 

 the bracts which subtend the heads and into the upper leaves. This series then represents a sort 

 of repetition of the character of the leaf in the calj-x and is noticed in various species in this genus. 

 The leaf-segments and calyx-lobes in Navarretia squarrosa are very similar or recall features one 

 of the other, as also in N. atractyloides. In nearly all the calyx variations of N. cotulaefolia cited 

 above there are likely to be only four calyx-lobes instead of five, the corolla being concurrently 

 tetramerous. 



Superficial characters used in differentiating Navarretia species are often inconstant or un- 

 satisfactory, but in some cases such differentiae may be strengthened by attention to the anatomy 

 or a more rigid analysis of floral or carpological organs, even to the kind of variation in an organ 

 and its degree of variation. 



A. Capsule not regularly dehiscent, the walls thin and transparent and closely covering the 



seeds which are agglutinated into a mass; seeds favosely pitted; 



flowers white or pale blue. 



1. Style 2-cleft or entire ; stamens exserted. 



Heads surrounded by foliaceous bracts (usually much reduced in no. 2) ; calyces glabrous (or 



rarely thinly puberulent) but with a ring of hairs on the inside of tube just below the 



sinuses ; stamens inserted about middle of corolla-throat ; style not cleft or slightly 2-cIef t. 



Stems erect or spreading (low and much-branched in no. 2) ; calyx usually glabrous outside. 



Heads % to 1 inch across, the outer foliaceous bracts conspicuously exceeding the heads; 



calyx-lobes entire, shorter than the corolla; Great Valley and North Coast 



Eanges 1. N. leucocephal/i. 



Heads small (3 to 6 lines across), dense, the bracts scarcely or not at all exceeding the 

 heads ; calyx-lobes entire or bifid or trifid, about as long as or slightly exceeding 



the corolla; n. Sierra Nevada and North Coast Ranges 2. A'', minima. 



Main stem short so that the primary flower cluster is seated on the ground, the branches 

 radiating from beneath it and prostrate ; calyx puberulent outside, its lobes entire 



or 2 of them bifid or trifid ; Monterey Co. to coastal S. Cal 3. N. prostrata. 



Heads subtended by rigidly acerose-toothed bracts, these and calyces densely white-tomentose (or 

 becoming glabrate in var. propinqua) ; stamens inserted near base of corolla-throat ; style 

 2-cleft; widely distributed 4. N. intertexta. 



S. Style 3-cleft. 



Stamens exserted, inserted on base of corolla-throat; bracts and calyces pubescent; ividely dis- 

 tributed 5. N. tagetina. 



Stamens included, inserted about middle of corolla-throat on very short filaments ; bracts and 

 calyces nearly glabrous, the margins of the bracts ciliate and the calyx with a ring of 

 hairs on the throat just below sinuses; n. Cal., rare 6. JV. mthuligcra. 



B. Capsule regularly dehiscent by valves and releasing the seeds 

 which are free from each other. 



1. Capsule 1-celled (sometimes imperfectly 2-celled) ; stamens 

 exserted; style 2-cleft (or sometimes entire). 



Capsule dehiscent from the base, 4 (or 8)-valved; leaves and bracts bipinnatifid, sometimes with 

 an oblaneeolate spatulate-dilated terminal segment. 

 Stamens inserted about middle of corolla-throat. 



Capsule 4 to 8-seeded; flowers 5-merous; corolla yellow with 5 dark purple spots in the 



throat; Sierra Nevada foothills. Great Valley and South Coast Eanges 



7. N. nigellaeformis. 

 Capsule 1 or 2-seeded ; flowers 4 or 5-merous. 



Heads villous and commonly minutely glandular, the body of the bracts and the 

 calyx-tubes glabrous or nearly so ; corolla creamy or dull white ; Coast 



Ranges and Sacramento Valley 8. N. cotulaefolia. 



Heads white-tomentose or densely white-villous; corolla blue, the throat white with 

 a purple spot at the base of each lobe ; n. Cal 9. N. bowmanae. 



