162 POLEMONIACEAE 



basal pair, or again the leaves may be quite entire. A conspicuous feature is the fascicles of short 

 leaves in the axils along the stems. This form of the coast line corresponds fairly well to a speci- 

 men collected by David Douglas, a part of the original Douglas collection of Hugelia densifolia 

 made in California (Herb. Univ. Cal., the contemporaneous label bearing the citation "Bot. Eeg. 

 t. 1622" and "Hort. Soc. 1833"). Douglas was sent to California by the London Horticultural 

 Society. He journeyed by land from Monterey to Los Angeles and although he doubtless made 

 an expedition to the inner Coast Eange, it is probable, indeed practically certain, that the type of 

 Hugelia densifolia was collected by him along the coast line. Coast line collections of this typical 

 form from San Luis Obispo Co. to Santa Barbara Co. may be cited: Morro, Hoadliouse ; Santa 

 Maria, Bloclnnan; Lompoc, Hoffmann. In the Blochman collection (as occasionally in this coast 

 line form), flowers or flower heads are developed on short branehlets arising from the buds of the 

 axillary fascicles (thus simulating H. virgata). There is, however, in coast liue forms every shade 

 of intergrade from the form with the leafy fascicles just described to a form with weak buds 

 in the leaf -axils or mth naked axils. The form with weak axillary fascicles corresponds to the 

 Douglas collection described by Bentham under the name Hugelia elongata, as evidenced by a 

 Douglas collection (Univ. Cal. Herb., contemporaneous label, with citation of "Bot. Eeg. 1622" 

 and "Hort. Soc. 1833"). This form is in reality a part of a series and is in this sense indistin- 

 guishable from the coast line form with axillary leaf fascicles and occasional small lateral heads 

 as noted above, which chances to represent the botanical or formal type. The presence of lateral 

 as well as terminal flower clusters is not significant, since such a condition may break out at various 

 interior stations of the "elongata" form (Banning, Gilman 746) ; such plants differ only slightly 

 from the "elongata" form. Moreover, a terminal head is sometimes replaced by a corymbose 

 cluster of small heads which approximates in form the development of lateral heads from lateral 

 leafy buds. In the "elongata" form 1 or 2 pairs of lobes are usually set below the middle of the 

 leaf -rachis or sub-basally set, with the terminal lobe or rachis much elongated, entire and gladiate. 

 The range of the species may be validated as follows. South Coast Eanges: mts. of Santa Clara 

 Co. (Greene, Man. Eeg. S. F. Bay 249) ; "Vancouver Pinnacles (6 mi. n.), San Benito Co., Howell 

 11,524; Tassajara road, Santa Lucia Mts., Hall 10,077; Callender, San Luis Obispo Co., Cooper 

 157; Estrella, San Luis Obispo Co., Jared. Cismontane S. Cal.: Casmalia, Santa Barbara Co., 

 Cooper 244; Big Pine Mt., San Eafael Mts., Hoffmann; Mono Flat, upper Santa Inez Eiver, A. L. 

 Grant 1688; Cuddy Canon, Mt. Piiios, H. # M. Hearing 1383; Aliso Canon, Santa Monica Mts., 

 Barber 192; Mt. Wilson, Peirson 521; Mt. San Antonio, San Gabriel Mts., Abrams 2682; San 

 Bernardino, S. B. # W. F. Parish; Santa Ana Eiver near Eedlands, Parish; Banning, If. F. Gilman 

 746 ; Saunder's Mdw., San Jacinto Mts., C. V. Meyer 622 ; Hemet Valley, San Jacinto Mts., Orcutt ; 

 Laguna Mts., Cleveland ; Jacumba, San Diego Co., Cleveland. Eanges on south side Mohave Des- 

 ert: Black Eock, near Warrens Well, Epling 4" Sobison; Eock Creek, n. slope San Gabriel Mts., 

 Peirson 482. Southern Sierra Nevada: Kern Co. (Greenhorn Summit, C. N. Smith 35; Pahute 

 Peak, Purpus 5087 ; Onyx, Voegelin) ; Inyo Co. (Bishop, Almeda Nordyke; Kcarsage Mill, Jepson 

 902 ; Independence, Aliee Rhine; Hunters Flat, Otto Renner 66; Cottonwood Creek, near Tknosea 

 Peak, Jepson 5091). 



Within the range of the species occur three phases described as varieties. All of these inter- 

 grade numerously to the usual form but each has some geographic significance, and intergradation 

 is mainly on the borders and not ■within the limits of the most marked areal development in 

 each case. 



Var. mohavensis (Craig) Jepson comb. n. Plants low (3 to 11 inches high) ; herbage white- 

 lanate; leaves % to 1 (or 2) inches long, with 1 to 3 pairs of tooth-like lobes; heads many to 

 numerous. — Sandy flats or sand dunes, 2000 to 4000 feet: Inyo Co. (Independence) ; Mohave Des- 

 ert (Kelso; Eedrock Canon; Muroc; Lancaster, — all ace. Bull. Torr. Club 64:392-3). 



Var. sanctanun (Mlkn.) Jepson. Herbage and heads densely white-woolly; corolla longer 

 (1 to 1% inches long). — San Bernardino Valley (San Bernardino, Parish 4178; MUl Creek, Lem- 

 mon; Santa Ana Eiver betw. Eedlands and Highlands, F. M. Eeed 3107). 



Var. austromontana (Craig) Jepson comb. n. Stems rather densely leafy; herbage mostly 

 subglabrate; leaves mostly shorter than the prevailing form (% to 1 inch long), the 2 to 4 pairs 

 of lobes equally spaced on the rachis as a rule, the terminal lobe thus short. — Montane, 4000 to 

 6000 feet: Lake Arrowhead, San Bernardino Mts., Braunton 1065; San Jacinto Mts., Gilman 

 747 ; Dos Palmos Spr., near Pinyon Flat, Santa Eosa Mts., Mum 15,105 ; Palomar Mt., Chandler 

 5372; Laguna Mts., Cleveland. 



Though there is, in a sense, so much of variation in this species group, it should be noted that 

 variational differences have here been emphasized. The features common to this species are 

 marked — so well marked as to unite the entire representation of it in one species unit with dis- 

 tinctive characteristics. It is seldom, indeed almost never, that a misdetermination is found in 

 collections of Hugelia densifolia. 



Eefs.— Hugelia densifolia Benth., Bot. Eeg. sub t. 1622 (1833), type from Cal., Houalas; 

 Jepson, Man. 792 (1925). Giliu densifolia Benth.; DC, Prodr. 9:311 (1845) ; Jepson, Fl. W. 

 Mid. Cal. 427 (1901), cd. 2, 332 (1911). Navarretia densifolia Brand; Engler, Pflzr. 4==0:165 

 (1907). Gilia hugelia Steud., Nom. Bot. ed. 2, 1:683 (1840). H. elongata Benth., Bot. Eeg. sub 

 t. 1622 (1833), type from Cal., Douglas. Gilia elongata Steud., I.e. G. densifolia var. elongata 



