166 POLEMOXIACEAE 



197; Bubbs Creek, Kings River Canon (Sierra Club Publ. 27:70) ; Mono Mdw., South Pork San 

 Joaquin River. A. L. Grant 1400; Fales Hot Sprs., Mono Co., Ottlry 1027; Lakeside Park, Lake 

 Tahoe, Geis 107; Sierra Valley, e. Sierra Co., Lemmon; Beckwith Pass, Lassen Co., Jepson 7757; 

 Eaglo Lake, Baker 4' Nutting. North Coast Ranges: betw. Burns Valley and Borax Lake, Lake 

 Co., Hoover 3553; Hy.ampum, Trinity Co., Chesnut 4' Drew; Hay Fork Valley, Trinity Co., Tracy 

 6463. Western Nevada on the easterly slope of, or in areas bordering, the eastern Sierra Nevada: 

 betw. Washoe and Franktown, Heller 10,603a; Eagle Valley, Ormsby Co., C. F. Baler 1403. 



Var. floccosa (Gray) Jepson comb. n. Stem erect, paniculately branched, sometimes dif- 

 fuse, sometimes subsimple, 4 to 11 inches high; herbage floceose-tomentose, more or less glabrate; 

 leaves narrowly linear, % to 1 inch long, with 1 (or 2) pairs of linear lobes at or near the base; 

 bracts palmately 3 or 5-cleft, the middle lobe elongated; heads persistently woolly, mostly 2 to 

 6-flowered, the body ovate or subglobose, (2 or) 4 to 7 lines broad; calyx equaling or a little 

 exceeding corolla-tube; corolla blue or white or pale lilac, 4 to 7 lines long, the throat yellowish 

 ■with maroon markings; corolla-tube and -throat 1% to 2% times as long as the calyx, sometimes 

 barely exceeding the calyx ; stamens inserted about middle of corolla-throat or a little below, the 

 anthers exserted from throat. — Dry flats or sandy hills, 2000 to 5000 feet: east side or easterly 

 valleys of the Sierra Nevada from Inyo Co. to Lassen Co.; inner South Coast Range in Fresno and 

 San Luis Obispo Cos. ; inner North Coast Range in Lake Co. East to Utah and north to Idaho and 

 eastern Washington. June-Aug. 



Tax. note. — The name Hugelia filifolia var. floccosa (Gray) Jepson rests on Gilla floccosa 

 Gray. Since the name used by this author is only of varietal rank, it is not necessary under 

 the International Rules to establish valid publication for the name Gilia floccosa Gray as applied 

 to the Great Basin plant, but it is of scientific interest to do so. In publishing Gilia floccosa it 

 is clear that Gray did not base his proposed species on "Hugelia floccosa Nutt. in herb.," which was 

 a nomen nudum and cited merely as a possible synonym, and as a possible synonym only because 

 it was to Gray an uncertainty. Gray's description of his species and indication of its geographic 

 range applies mainly if not wholly to transmontane plants. Moreover, a new name with a properly 

 published description cannot be invalidated because of wrongly citing the name of a different and 

 valid species as a synonym. In this case Gray's name cannot be invalidated by the name Hugelia 

 lutea Bcnth., which belongs to a different species of the California coast line. Moreover, although 

 Hugelia lutea was cited by Gray as a synonym, it is so cited with slightly qualifying phrases which 

 indicate shadows of doubt. Yet again, further light on Gray's main intent may be had from sub- 

 sequent works written at the Gray Herbarium, namely: Botany of the King Expedition (207, — 

 1871, in which application of the name, Gilia floccosa, is limited wholly to Great Basin plants) ; 

 Botany of the California Geological Survey (1:495, — 1876, in which the name is applied primarily 

 to transmontane plants); Synoptical Flora (2:143, — 1878, in which the name refers solely to 

 Great Basin or interior plants, the name Hugelia lutea Benth. being definitely excluded). All 

 these works, thus, make Gilia floccosa Gray a Great Basin plant. Gilia floccosa Gray, a transmon- 

 tane plant, was sufficiently published. One cannot disqualify this name by reason of Hugelia 

 floccosa Nutt., which had never been published and which was cited only with uncertainty as a 

 nomen nudum; nor by reason of the name Hugelia lutea Benth., which belongs rightly to another 

 species. Since, therefore, the name Gilia floccosa Gray is applicable only to the interior plant and 

 not to a coastal plant, the following later names of the interior plant are added to the synonymy: 

 GUia wilcoxii Nels., Navarretia wilcoxii Brand and Welwitschia wilcoxii Eydb. 



Field note. — The range of variation in plant habit in Hugelia filifolia var. floccosa is rather 

 marked. Slender tall forms and coarse stocky forms, otherwise identical, may be found in the 

 same collection. Densely branched forms are very unlike, in appearance, slender erect few- 

 branched forms. Heads in different collections vary in size, the forms with large crowded several- 

 flowered heads differing noticeably from forms with narrow few-flowered heads, these latter 

 simulating closely the aspect of typical Hugelia filifolia. 



Locs. — East side of the Sierra Nevada: Darwin Mesa, Otto Eenjier 80; Owens Valley, Horn 

 2850; Bishop, Inyo Co., Almrda NordyTce; Black Caiion, White Mts., Duran 2681; Mono Craters, 

 Feirson 10,755; Freshwater Spr., near Mono Lake, Ottley 1100; Bloody Canon, Chesmit 4' Drew; 

 Dog Valley, e. Nevada Co., Jeyson 15,049; Long Valley, Lassen Co., Jepson ni)'2. Inner South 

 Coast Ranges in western Fresno Co. and in San Luis Obispo Co.: Waltham Creek, Diablo Range, 

 Easttvood 4' Hoivell 5835; Black Mt., Pozo Range, Giff'ord; (these last two collections match very 

 closely Duran 2681 from the White Mts. east of the Sierra Nevada, cited above). Inner North 

 Coast Range: Snow Mt., Lake Co., K. Brandegee. 



Refs. — Hugelia filifolia Jepson, Man. 792 (1925). Gilia filifolia Nutt., Proc. Acad. Phila. 

 4:11 (1848), type loc. near Santa Barbara, Gambel. Navarretia filifolia Brand; Engler, Pflzr. 

 4=='':167 (1907). Eriastrum filifolium Woot. & Sta., Contrib. U. S. Nat. Herb. 16:160 (1913). 

 Welwitschia filifolia Rydb., PI. Rocky Mts. 688 (1917). Gilia virgata var. filifolia MIkn., ITuiv. 

 Cal. Publ. Bot. 2:39 (1904). G. floccosa var. filifolia Nels. & Mcbr., Bot. Gaz. 61:35 (1916). 

 G. sparsiflora Eastw., Proc. Cal. Acad. ser. 3, 2:291 (1902), type loc. Kings River Canon, East- 

 wood. Navarretia filifolia subsp. sparsiflora Brand; Engler Pflzr. 4-'''*:167 (1907). Gilia fili- 

 folia var. sparsifiora Mcbr., Contrib. Gray Herb. n. s. 49:57 (1917). H. filifolia var. sparsiflora 

 Jepson, Man. 792 (1925). Var. floccosa Jepson. Gilia floccosa Gray, Proc. Am. Acad. 8:272 

 (1870), "California to Arizona, interior of Oregon, and Utah"; of the various collections be- 



