GILIA FAMILY 



185 



Saiid dunes and beaches along the coast line, 5 to 50 feet : Humboldt Co. to San 

 Francisco Bay and south to Santa Barbara Co. ; also on an arm of Suisun Bay in 

 the lower San Joaquin Valley. May-June. 



Locs. — Betw. Trinidad and Little River, Humboldt Co., Tracy 2583; Alton, Humboldt Co., 

 Tracy 3672; Bodega Pt., Sonoma Co., Cooper 88; Bolinas Bay, K. Brandegee; Antioeh (relict 

 sand dune in interior but situated on an arm of Suisun Bay), Swenson; Presidio, San Francisco, 

 Jepson 15,081; Lake Merced, San Francisco, K. Brandegee ; Antonio siding betw. Casmalia and 

 Surf, Santa Barbara Co., K. Brandegee. 



Geog. note. — In its best developed or most typical form, perhaps always, Gilia chamissonis is 

 an inhabitant of pure sand on ocean dunes. This fact, in connection with its morphological char- 

 acters, furnishes the basis for its recognition as a species. It does, however, in reality or in appear- 

 ance intergrade to Gilia staminea in the South Coast 

 Ranges, while northern plants of it simulate Gilia 

 capitata in certain particulars. Such evidence tends 

 to argue that it should be disposed as a variety of Gilia 

 staminea or Gilia capitata. If the principle of even 

 moderate intergradation be enforced too strongly 

 against specific validity, the fall of Gilia chamissonis 

 would also drag down Gilia capitata, G. staminea, G. 

 achilleaefolia, G. multicaulis and various other species 

 in this genus. While Gilia chamissonis is, admittedly, 

 kept as a species with considerable diffidence, it is re- 

 tained, partly by reason of its geographic consistency. 



Var. regina Jepson comb. n. Stem once or twice 

 forked, arising from a conspicuous basal tuft of 

 leaves ; leaf -segments broader, the rachis somewhat 

 woolly ; peduncles long, stout and naked ; heads large 

 (1 to 1% inches wide) ; corolla large (4% to 5 lines 

 long, 3 to 6 lines wide) ; anthers large. — Sand dunes. 

 Point Reyes peninsula, Marin coast (L. Tinsley). 



Refs. — GuiA CHAMISSONIS Greene, Erythea 3 : 

 105 (1895), type loc. "sand-hills of San Francisco," 

 Greene, based on the same characteristic plant of the 

 San Francisco sandhills as Polemonium capitatum 

 Esch., Mem. Acad. Petrop. 10:282 (1826), "in Novae 

 Californiae arenosis" [San Francisco], Eschscholtz ; 

 not Gilia capitata Dougl. (1826). G. achilleaefolia 

 subsp. chamissonis Brand; Engler, Pflzr. 4-^": 111 

 (1907), in part. G. achilleaefolia var. chamissonis 

 Nels. & Mcbr., Bot. Gaz. 61:34 (1916). G. achilleae- 

 folia var. tmnentosa Eastw. ; Brand in Engler, Pflzr. 

 4"'':111 (1907), type loc. Bodega Pt., Sonoma Co., 

 Eastwood 2. Var. eeoina Jepson. G. capitata var. 

 regina Jepson, Man. 795 (1925), type loc. Pt. Reyes 

 peninsula, Jepson 8315. 



Fig. 384. Gilia achilleaefolia Benth. 

 a, lower part of plant, X % ; 6, infl., X 

 % ; c, fl., X 2%. Drawn from Douglas 

 type. Royal Botanic Gardens, Kew (Herb. 

 Benth.). 



11. G. achilleaefolia Benth. (Fig. 384.) Stem erect, erectly branching or 

 simple, 7 to 17 inches high ; herbage glabrous to puberulent ; leaves 1 to 5 inches 

 long, the blades pinnately or mostly bipinnately divided into linear segments; 

 flowers about 5 to 15 in rather dense terminal cymes, the cymes capitate but not 

 globose; calyx puberulent, 3 to 3i/4 lines long, its triangular-acute or lanceolate 

 teeth 1/2 as long as the tube; corolla violet-blue or rarely white, broadly funnelform, 

 5 to 81/^ lines long, 21/0 to 4 times as long as the calyx; corolla-throat and -limb ex- 

 ceeding (sometimes much exceeding) the corolla-tube, the throat extremely ample 

 and the lobes ovate to oblong; corolla-tube much shorter than, equaling or only 

 slightly exceeding the calyx; stamens included in corolla-throat; style usually 

 exserted. 



Rocky foothills and fiats, 100 to 1200 feet: mostly along or near the coa.st line 

 from Santa Clara Co. to Los Angeles Co. Mar.-May. 



Tax. note. — The true Gilia achilleaefolia Benth., as here described, appears to be the plant 

 on or near the coast line from middle California to Los Ajigeles Co., that is, the species described 

 as Gilia abrotanifolia Nutt. by Greene. It is remarkable for its broad or turbinatcly expanded 

 corolla-throat which usually exceeds the short narrow tube. The inflorescence is a much con- 



