186 



POLEMONIACEAE 



densed or capitate cyme with the flowers all erect; the calyces are villous or puberulent ; the sta- 

 mens are included. The species here described as Gilia staminea Greene, that is Gilia aehilleaef olia 

 of Gray (Syn. Fl. 2:147,-1876) in great part, Jepsou, Fl. W. Mid. Cal. 426 (1901) and many 

 other authors, is a plant which often simulates the true Gilia achilleaefolia in habit and in flowers 

 very closely ; its inflorescence is likewise a dense head-like cyme, but its heads, whether symmetric- 

 ally or irregularly globose, bear the flowers spreading (all around) outwards; the stamens are 

 exserted; the calyces are densely woolly to glabrate; it ranges widely in the ulterior, although also 

 occurring along the coast. In the mass, the technical differences between the two species are 

 slight in degree and are apparently degraded by intermediates. The decision here made that Gilia 

 abrotanifolia Nutt. equals Gilia achilleaefolia Benth. is based first upon Douglas specimens now 

 in the University of California Herbarium which are presumably part of tlie original Douglas 

 collection of Gilia achilleaefolia made in California; second, upon the illustration of Gilia achil- 

 leaefolia in the Botanical Eegister (t. 1682) ; and third, upon 

 a drawing of the type specimen made under the direction of 

 the author in the Herbarium Benthamianum. That the Bo- 

 tanical Register illustration represents the south coast line 

 plant heretofore called Gilia abrotanifolia Nutt. is noted by 

 R. F. Hoover. It is not the more widely dispersed plant hith- 

 erto called G. achilleaefolia in American floras which latter is 

 now described as Gilia staminea Greene. 



Gilia achilleaefolia Benth. is also closely allied to Gilia 

 multicaulis Benth. Both are coastal species, but GUia multi- 

 caulis has a more extensive longitudinal range and extends 

 further to the interior. In the northern and southern parts of 

 its range Gilia achilleaefolia intergrades with Gilia multi- 

 caulis, but has its extreme expression in coastal San Luis 

 Obispo County where, apparently, typical Gilia multicaulis is 

 less common. 



Collections of Gilia achilleaefolia are cited as follows : Mt. 

 Hamilton Range (w. slope) : Alum Eock, Pendleton 6S4; 

 Coyote Caiion (mouth), e. of Madrone, Hoover 3268. Santa 

 Cruz Mts.: Stanford (foothills near), C. F. Baker 643; Los 

 Gates (foothills w.). Heller 7396. Gabilan Range: San Juan, 

 San Benito Co., Elmer 5037. San Luis Obispo Co.: Paso 

 Robles, Barber; San Luis Obispo, M. M. Miles 134; Arroyo 

 Grande, Alice King. Coastal S. Cal.: Gaviota Pass, Santa 

 Barbara Co., Brewer 390 ; Saugus, Los Angeles Co., Geo. B. 

 Grant 5433 ; Claremont, Chandler. Desert slope San Bernar- 

 dino Mts.: headwaters Mohave River, Palmer 411. 



Refs. — Gilia achille.\efolia Benth., Bot. Reg. sub t. 

 1622 (1833), type from Cal., Douglas; Lindley, Bot. Reg. t. 

 1682 (1835) ; not of Gray, Jepson and other American authors. 

 G. abrotanifolia Nutt.; Greene, Erythea 3:104 (1895), type 

 loc. Santa Inez Mts., back of Santa Barbara, Nnttall ; (since 

 the binomial is Nuttall's, it seems better to designate Nuttall's 

 specimen as the type of Greene's description, although it is not 

 mentioned first by Greene). G. achilleaefolia subsp. abrotani- 

 folia Brand; Engler, Pflzr. 4-^'':lll (1907), excluding reference to Mariposa Co. G. multicaulis 

 var. eximia Mlkn., Univ. Cal. Publ. Bot. 2:35 (1904), type loc. Evergreen, Santa Clara Co., Davy 

 1883. G. muUicaxdis subsp. eximia Brand, I.e. 110. 



12. G. multicaulis Benth. (Fig. 385.) Stem simple to bushy-branching, strict 

 to ascending, 6 to 20 inches higli; herbage glabrate or puberulent, the inflorescence 

 puberulent or with short gland-tipped hairs; leaves chiefly basal or sub-basal, 

 slightly reduced up the stem, the blades never glandular or rarely so, pinnate or 

 bipinnately divided, the lobes filiform or linear, remote or crowded, 1^/2 to 5 (or 7) 

 lines long ; lower leaves 1 to 4 inches long, petioled, upper ones sessile ; flowers 2 to 

 7 in terminal glomernles, or sometimes solitaiy ; calj^x cleft %, its lobes triangular- 

 lanceolate, cuspidate, the scarious intervals often purple ; corolla funnelform, whit- 

 ish or smoky-blue or pinkish, Sy^ to 5 (usually aboiit 4) lines long, about twice as 

 long as calyx, its tube yellowish ; stamens shorter than the corolla-lobes, inserted 

 at the sinuses ; capsule ovate-oblong, the cells many-seeded ; seeds showing spiricles 

 when wetted. 



Dry rocky or gravelly hills and flats, 175 to 3000 feet : Marin Co. ; South Coast 

 Ranges; cismontane Southern California; west side of the Colorado Desert; south 

 to Lower California. Apr.-June. 



Fig. 385. Gilia multicaulis 

 Benth. a, habit X % ; b, fl., X 3. 

 DrawTi from Douglas type. Royal 

 Botanic Gardens, Kew (Herb. 

 Benth.). 



