GILIA FAMILY 187 



Tax. note. — Gilia multicaulis, based on a Douglas plant from California, was, almost -n-ith- 

 out doubt, discovered in the South Coast Eanges. The type (preserved at Kew; see fig. 38i) is a 

 plant 8 inches high with several erect stems from the base and the flowers borne in terminal 2 to 

 7-flowered glomerules. The stems are somewhat leafy and a few pinnately divided leaves are 

 borne at base. The calyx and pedicels are provided with short gland-tipped hairs. The calyx- 

 lobes are about % as long as the tube. The throat of the funnelform corolla is shorter than the 

 corolla-tube. On the one hand Gilia multicaulis approaches so closely to Gilia achilleaefolia that 

 some collections seem as well referred to one species as to the other. On another hand, through a 

 chain of simulations, it exhibits the habit and the corolla markings of Gilia tricolor. Corollas so 

 marked are, however, quite small and present a sufficient contrast to the ample corollas of Gilia 

 tricolor. 



In its prevailing form Gilia multicaulis is a leafy-stemmed plant without marked basal leaf- 

 rosette, its flowers borne in few- to several-flowered glomerules. In its inflorescence it is extremely 

 variable and solitary flowers or loosely flowered forms are frequent. Like most highly plastic 

 Gilia species it has been much subject to specific segregation by some botanists. A few of its 

 extreme phases are described below as subspecies or varieties. The following collections arc cited 

 as validating the range of the species: 



Loes. — Marin Co.: Mt. Tamalpais, Jcpson 7556. South Coast Eanges: Berkeley Hills, Jepson 

 15,072; Mt. Diablo (Erythea 1:176) ; Mt. Day, Mt. Hamilton Eange, K. J. Smith; Ben Lomond, 

 Santa Cruz Mts., K. Brandegee; Carmel Mission, Heller 6525; Cholame, San Luis Obispo Co., 

 Wiggins 5778. Cismontane S. Cal.: Santa Rosa Isl. and Santa Cruz Isl., T. Brandegee ; Santa 

 Maria (7 mi. se.), Jepson 19,098; Tepusquet Canon, near Sisquoc Eiver, n. Santa Barbara Co., 

 M. 4" ^- Bearing 1430 ; Big Tujunga Caiion, San Gabriel Mts.. Peirson 157a ; Saugus, Davy : Eaton 

 Cation, San Gabriel Mts., Peirson 424; Henninger Flat, San Gabriel Mts., Peirson 157; San Ber- 

 nardino, Parish; Banning, Clokeif 4" Templeton 4661; Anaheim plains, Alice King; Eiverside, 

 Jepson 1230; Elsinore, Munz 12,024; Oak Grove, ne. San Diego Co., Jepson 17,149; San Pasqual, 

 San Diego Co., Jepson 19,176; Viejas Valley, San Diego Co., Jepson 11,836; Mesa Grande, San 

 Diego Co., E. Fergxison 39; La Junta, near Jamul, San Diego Co., Newlon 342. West side Colo- 

 rado Desert: Andreas Canon, San Jacinto Mts., Newlon 433b. 



Var. clivormn Jepson var. n. Stem branched or sometimes simple, 4 to 15 inches high; herb- 

 age densely glandular to glabrate; leaves 6 to 12 lines long, pinnately or bipinnately divided into 

 very narrow rather remote lobes, the upper leaves sessile, the lower petiolate ; flowers in terminal 

 few-flowered glomerules; calyx cylindric, cleft %, the hyaline intervals extremely narrow and 

 typically dark purple; corolla 3 to 4 lines long, l^fj to 2 times as long as calj^, dull blue, the throat 

 ornamented with 5 dark purple spots ; corolla-lobes narrowly oblong to broadly ovate, little spread- 

 ing in anthesis; stamens inserted at sinuses, shorter than corolla-lobes; capsule ovate, 2 to 2% 

 lines long, the cells many-seeded; seeds with a narrow membranous margin. — (Caulis ramosus vel 

 subsimplex, 4—15 une. altus; herba dense glandulosa vel glabrata; folia 6-12 lin. longa, uni- vel 

 bi-pinnatifida, lobis angustis subremotisque; glomeruli terminales pauciflores; calyx cylindricus, 

 intervallis hyalinis angustissimis atropurpureisque; corolla 3-4 lin. longa, caerulea remissa, calyce 

 1%-2-plo longior, fauce maculis 5 atropurpureis ornata; corollae lobi anguste oblongi vel late 

 ovati ; stamina sinibus affixa, corollae lobis breviora; capsulae loculi polyspermi; semina anguste 

 membranacea alata.) — Gravelly hillslopes or canon sides, 20 to 1200 feet: South Coast Eanges 

 from Contra Costa and San Mateo Cos. to San Luis Obispo Co.; northwestern Santa Barbara Co. 

 Mar.-May. Here are included plants of the South Coast Eange hills back of the coast line which 

 previously have been named Gilia millefoliata or Gilia multicaulis var. millefoliata. The true 

 Gilia millefoliata P. & M. is, however, a plant of the coast line sand dunes from Sonoma Co. to 

 Del Norte Co., which is here referred to Gilia inconspicua Sweet and is held different from Gilia 

 multicaulis var. clivorum of the South Coast Eange hills. "Var. clivorum is a slender plant with 

 smaller corollas; its calyx is not very markedly accrescent; its capsules are oblong-ovate and 2 

 lines long. The stems of Gilia inconspicua by contrast are a little stoutish and rigid ; the calyx 

 is strikingly accrescent; the oblong capsules are 3 to 4 lines long. There is also marked geogra- 

 phic segregation. The following are citations of var. clivorum: 



Locs. — Mitchell Caiion, Mt. Diablo, C. F. Balcer 2951; Crystal Springs Lake, San Mateo Co., 

 C. F. Baker 467; San Martin, Santa Clara Co., Chandler 897; Hayes sta., s. end Panoche Hills, w. 

 Fresno Co., Jepson 16,980; Paso Robles, Barber (type) ; San Luis Mt., Summers. Santa Barbara 

 Co.: San Miguelito Canon, Lompoc, Mum 10,249. 



Var. millefoUa Gray. Stem leafy; leaves Va to 1% inches long, thrice pinnate with numerous 

 short linear or subfiliform segments, the segments crowded, % to 1% lines long; calyx-teeth 

 spreading; corolla bluish, very narrowly funnelform, its tube more slender, the throat scarcely 

 expanded, 3% to 4 lines long; capsule long-oblong. — Sant-i Barbara Isls. ; Santa Cruz Isl., T. Bran- 

 degee ; Green Caiion, Santa Rosa Isl., Hoffmann; San Clemente Isl., Mum 6633. South to Gua- 

 dalupe Isl., Anthony 235. 



Var. peduncularis (Eastw.) Jepson comb. n. Stem slender, erect and sparsely branched, 8 to 

 16 inches high, sometimes diffusely branched from the base; herbage subglabroiis or thinly and 

 finely villous, the peduncles or pedicels minutely glandular-puberulent ; leaves % to 2% inches 

 long, the blades pinnately divided into 3 to 7 (or 9) somewhat remote segments, the segments 



