188 pou:moniaceae 



linear or narrow-oblanecolate, 2 to 7 lines long, entire (or rarely mth 1 prominent lobe) ; flowers 

 solitary on filiform naked pedicels 1^ to 2 inches long, or sometimes some of the flowers in clusters 

 of 2 or 3 on very short pedicels and the clusters on long naked peduncles ; calyx puberulent, % to 

 % as long as the corolla, its lanceolate teeth about equaling the calyx-tube ; corolla blue or whitish, 

 tubular-f unnelform, 3 to 5 lines long, its tube included in the calyx-tube or barely exserted. — Hill- 

 slopes, 500 to 2100 feet: Vaca Mts. ; South Coast Ranges from Contra Costa Co. to San Luis Obispo 

 Co. and western Fresno Co.; Greenhorn Mts., Kern Co.; coastal Southern California (rare) ; north- 

 western Colorado Desert. South to Lower California. Apr. 



Geog. note. — Gilia multicaulis var. peduncularis seems little more than a loose form of Gilia 

 multicaulis but is here given varietal status because of the characters developed in its extreme 

 phase : the slender habit, the few long remote segments of the pinnately divided leaves, the long 

 naked filiform pedicels, the frequently solitary flowers and the small corollas. This varietal group, 

 however, as here defined, does not seem to represent a natural entity of a single genetic origin 

 since it occurs sporadically throughout the range of the species. The case against recognition of 

 Gilia peduncularis as a species may be stated thus: Its aspect indicates it as a merely tenuous form 

 of Gilia multicaulis. Typically the stems are remotely leafy and the basal cluster of leaves weak, 

 but when the basal leaf-cluster is strongly developed, the cauline leaves are often much reduced 

 and the plant tends again to suggest the species. Although the flowers are solitary in the type 

 of Gilia peduncularis, collections often show flowers in glomerules (as in Gilia multicaulis) as 

 well as solitary. A plant with pcdicelled flowers may often be found in a collection consisting of 

 plants displaying quite glomerate flowers (that is, with 4, 5 or 6 flowers in a cluster), or often 

 several of the flowers of a glomerate plant are solitary. Furthermore, those plants whose flowers 

 are solitary rather than glomerate, or wholly solitary, differ so greatly among themselves that it 

 seems unnatural to segregate them as a species or perhaps even as a variety. Nor are they segre- 

 gated geographically from Gilia multicaulis, since they occur, rather, throughout the geographic 

 area of Gilia multicaulis. To give these long-pedicelled plants specific rank would do violence to 

 the plant population of Gilia multicaulis, as a whole, as found in its natural range. To the fore- 

 going the following note may be added. "In Gilia tricolor two forms likewise occur, the typical 

 form in which the flowers are congested and var. longipedicellata with a loose inflorescence. To all 

 appearances var. peduncularis bears the same relation to Gilia multicaulis as does var. longi- 

 pedicellata to Gilia tricolor." — R. F. Hoover. 



Locs. — Vaca Mts.: Dutton Canon, Jcpxon 15,068; Graveyard Hill, se. of Vacaville, Jepson 

 15,071 (grading towards var. clivorum). South Coast Ranges: Mt. Diablo, Bowerman 2839; 

 Kings Mt., San Mateo Co., Kcclc 2931; Mt. Hamilton Range (foothills e. of Gilroy), Hoover 

 3272; Charmichael Mill, Santa Cruz Mts., Pendleton 935; Sur River, Monterey Coast, Eastwood ; 

 Tassajara Hot Sprs., Santa Lucia Mts., Ferris 8320 ; Zapato Canon, Diablo Range, sw. Fresno Co., 

 Jepson 15,376 (flowers sometimes in compact clusters, thus approaching typical Gilia multicaulis). 

 Greenhorn Mts., Kern Co.: Poso Mine (3 mi. sw.), Poso Creek, C. N. Smith 50; Kern River Park, 

 Kranifs. Coastal S. Cal.: Zaca road, Santa Ynez Valley, Hoffniann : Refugio Pass, Santa Ynez 

 Mts., Hoffmann; Prisoners Harbor, Santa Cruz Isl., Hoffmann 794; Ramona, San Diego Co., 

 K. Brandcgce. Northwestern Colorado Desert: Andreas Canon, San Jacinto Mts., Newlon 432 

 (grading towards the species). 



Rets. — GiMA MULTICAULIS Benth., Bot. Reg. sub 1. 1622 (1833), type from Cal., Douplas (typ. 

 vidi) ; Jepson, Fl. W. Mid. Cal. 425 (1901), ed. 2, 331 (1911), Man. 796 (1925). G. multicaulis 

 var. tenera Gray, Proc. Am. Acad. 8:278 (1870), type from "Cal."; very slender, peduncles long, 

 the terminal clusters usually 2 or 3-fiowered or reduced to one flower and thus transitional towards 

 var. peduncularis; of the collections named by Gray the following is selected as lectotype (Santa 

 Cruz Redwoods, Bolander 55; Gray Herb.) in order to define the relation to var. peduncularis. 

 G. multicaulis var. detonsa Gray; Mlkn., Univ. Cal. Publ. Bot. 2:35 (1904), type loc. Berkeley 

 Hills. G. multicaulis var. alba Mlkn., Univ. Cal. Publ. Bot. 2:35 (1904), type loc. Los Gatos, Santa 

 Clara Co., A. L. Jag gar. G. oreophila Greene, Baker's West American I'lants 1:9 (1902), C. F. 

 Baker 865 [foothills near Stanford, Santa Clara Co], nomen nudum. G. inconspicua var. oreophila 

 Brand; Engler, Pfizr. 4-''°: 105 (1913). Var. clivorum Jepson. Var. millepolia Gray in Wats., 

 Proc. Am. Acad. 11:118 (1876), type loc. Guadalupe Isl., Palmer. G. vevinii Gray, Syn. Fl. ed. 2, 

 2:411 (1886), type loc. San Clemente Isl., Nevin ^ Lyon. G. multicaulis var. nevinii Jepson, Man. 

 796 (1925). Var. peduncularis Jepson. G. peduncularis Eastw.; Mlkn., Univ. Cal. Publ. Bot. 

 2:34 (May, 1904), type loc. Dutard ranch, near the boundary line betw. Santa Barbara Co. and 

 San Luis Obispo Co. (probably w. of Santa Maria), Fastwood; Jepson, Man. 796 (1925). G. pe- 

 dunculata Eastw., Bot. Gaz. 37:446 (June, 1904), type loc. "Alamo Creek," near Santa Maria, 

 "Santa Barbara Co.," Eastwood (this Alamo Creek is probably in San Luis Obispo Co.). G. pe- 

 dunculata var. calycina Eastw. I.e. 447, type loc. betw. Huasna and Pozo, San Luis Obispo Co., 

 Eastwood. G. peduncularis var. typica subvar. calycina Brand; Engler, Pflzr. 4-'^'*:108 (1907). 

 G. pedunculaia var. minima Eastw., I.e. 447, type loc. Cuyama, Eastwood. G. peduncularis var. 

 minima Brand, I.e. 108. G. multicaulis var. stricta Brand, I.e. 110, the name based on G. stricta 

 Scheele, Llnnaea 21:755 (1848), a garden plant cultivated in Germany. 



13. G. inconspicua Sweet. Stem stoutish, 1 or several from the base, erect or 

 ascending, 5 to 10 inches high; herbage, peduncles and calyces glandular-puberu- 



