GILIA FAMILY 197 



5 (or 13) lobes or leaflets; lobes or leaflets entii-e, rarely toothed, commonly very 

 unequal; blades of upper leaves commonly trifid, often very different in shape from 

 the lower leaves; flowers on slender pedicels, arranged in open irregular cymes 

 or glomerate, minutely bracteate or conspicuously leafy-braeteate ; calyx very 

 glandular-hairy, searious below sinuses to base ; calyx-lobes erect, linear or subulate, 

 equaling the tube or longer; corolla tubular-funnelform, white, blue, purple or 

 rarely red-pink, 2i/2 to 5 lines long, 2 to 3 times as long as the calyx, the limb 1 to 

 11/4 lines broad; stamens unequal and unequally inserted on the corolla-throat 

 (about the middle or below), included in the throat or the longer with anthers 

 barelj^ exserted; style included or rarely exserted; capsule globose, its cells 1 (rarely 

 2) -seeded; seeds becoming mucilaginous when wetted. 



Dry sandy flats or hills or wooded canon sides, 2500 to 9000 feet: common 

 throughout the Coast Ranges (mostly towards the interior). Sierra Nevada and 

 cismontane Southern California, less frequent on the east side of the Sierra Nevada, 

 rare in the Sacramento Valley (100 to 500 feet) and absent from the San Joaquin 

 Valley, Mohave Desert (except west side) and Colorado Desert. South to Lower 

 California, east to western Nevada and north to southern Oregon. Mar.-Aug. 



Locs. — Coast Ranges: Sisson, Siskiyou Co., Jepson 15,061; Hyampum, Trinity Co., dies- 

 nut 4- Brew; Greasewood Hills, near Mt. Tom, w. Tehama Co., Jepson 15,067; Alder Sprs. (10 mi. 

 e.), n. Glenn Co., Heller 11,4.38; Mt. Konoeti, Lake Co., Jepson 15,059; Howell Mt., Napa Range, 

 Jepson 15,060; Mt. Diablo, Bowerman 1907; Saratoga (s. of), Santa Cruz Mts., Pendleton 130; 

 Lopez Caiion, San Luis Obispo Co., Unangst. Sacramento Valley: Sacramento, T. Brandegee. 

 Sierra Nevada: Jonesville, C op eland ; Pioneer road sta.. North Fork Yuba River, Jepson 16,799; 

 Bear Valley, Nevada Co., Jepson 15,062; Strawberry road sta., Eldorado Co., Jepson 16,886; 

 Myers sta., near Lake Tahoe, Ottley 940; SUver Valley, Alpine Co., Jepson 10,148; Cow Creek, 

 Tuolumne Co., Jepson 6514; Heteh-Hetehy, Jepson 3449; Lake Merced, Merced River, Jepson 

 3197 ; Fresno Big Trees, Madera Co., Jepson 15,973 ; Graveyard Mdw., near Silver Pass, Fresno 

 Co., A. L. Grant 1513b; Big Creek, Fresno Co., I. T. WalTier; Shaver Lake, Fresno Co., Jepson 

 16,097 ; betw. Tehipite Valley and Gnat Mdw., E. Ferguson 535 ; South Fork Middle Tule River, 

 Jepson 4872; Portuguese Mdw., s. Tulare Co., C. N. Smith 16. East side of the Sierra Nevada: 

 Forestdale, 3W. Modoc Co., M. S. BaTcer; Susanville, T. Brandegee; Sierra Valley, Lemnion; Mo- 

 gul, Mono Co., Yates 5310; Pleasant Caiion, Panamint Range, Hall ^ Chandler 6967. Western 

 Mohave Desert: Manzana, Antelope Valley, Davy 2516. Cismontane S. Cal.: Mt. Piiios, Hoff- 

 mann; Zaca road, San Rafael Mts., Hoffmann; Vandeventer Flat, Santa Rosa Mts., Jepson 1416; 

 Escondido, San Diego Co., C. V. Meyer 117; Cuyamaca Mts., Palmer 234. 



Note on variation. — Gilia gilioides is remarkably unstable; it is unstable in habit, in shape 

 of leaves and in form of inflorescence, — these are the features most subject to variation. A char- 

 acteristic leaf diversity as found on a single individual is as follows: leaves entire or toothed on 

 the lower part of the plant, leaves pinnately divided (with 3 to 5 or 9 or sometimes 13 lobes) at the 

 middle of the plant, and leaves trifid above. The segments of the leaves or leaflets vary greatly 

 in size and shape. Variation in habit, ranging from coarse plants to delicate plants, is equally 

 striking. Notwithstanding all this, variation is confined within certain limits and tends to charac- 

 terize or place its peculiar seal upon the entity. The character of the variation, in other words, 

 marks well the species, almost as well as an invariable constant. Collections are, therefore, usually 

 referred with certainty to this species. Only three varieties are here described: 



Var. volcanica (Brand) Jepson & Hoover comb. n. Corolla 6 to 10 lines long, 2 to 3 lines 

 broad, the tube purple, the lobes white or pale; stamens unequal, the longer conspicuously exserted, 

 the filaments unequally inserted, the upper at about middle of throat, the lower at base of throat; 

 style shorter than corolla-tube. — Sierra Nevada foothills from Eldorado Co. to Mariposa Co. ; also 

 in Lake Co. (intermediate towards the species). 



Locs. — Sierra Nevada foothills: Sweetwater Creek, Eldorado Co., K. Brandegee; Avery sta., 

 Calaveras Co., A. L. Grant; Bear Mt., Calaveras Co., Davy 1456; Yankee Hill, Tuolumne Co., 

 Jepson 6416; Columbia, Jepson 6418; Cold Spr., Bald Mt., Tuolumne Co., Jepson 6465; Coultor- 

 ville, Mariposa Co., Hoover 2160. Lake Co. (corolla similar, but stamens apparently never ex- 

 serted) : Snow Mt., T. Brandegee; Binkley ranch, near Cobb Mt., Jussel 159. 



Var. ianthina Jepson & Hoover var. n. Leaves or their lobes usually narrow, the lower leaves 

 (save in depauperate plants) mostly with 5 to 15 lobes ; flowers 3 to 7, crowded in capitate clusters; 

 corolla bright violet-blue. — (Folia vel lobi fere angusti; folia basilaria plerumque 5-15-lobata; 

 flores 3-7, dense capitati; corolla violaceo-azurea.) — Inner Coast Range and its cast-side foothills 

 from western Shasta Co. to southwestern Kern Co. ; Sierra Nevada foothills from Fresno Co. to 

 Kern Co. ; south to the western San Gabriel Mts. It intergrades to typical Gilia gilioides. 



Locs. — Inner Coast Range: Anderson, Shasta Co., Alice King; Paskenta (6 mi. w.), Virginia 

 Bailey; Lodoga, Colusa Co., Jepson 16,271 (type) ; Mt. Diablo, Bowerman 2919; Mt. Hamilton, 



