QILIA FAMILY 209 



Tax. note. — Linanthus liniflorus vs. Linanthus pharnaceoides. These two forms were first 

 published under Gilia by George Bentham. The descriptive terms used by Bentham in the original 

 diagnoses of Cilia liniflora and Gilia pharnaceoides (Bot. Reg. sub t. 1622), both collected by 

 Douglas in "California," are e.xactly the same, save that in the former the corolla is described as 

 thrice longer than the calyx, in the latter twice longer. In DeCandoUe's Prodromus (9:315), 

 Bentham adds that the throat is very pilose in G. liniflora and puberulent in G. pharnaceoides. 

 Since the types were collected by Douglas in California it would seem that both forms must have 

 been discovered in the coastal region, almost certainly in the South Coast Eanges, which were 

 traversed by Douglas; but in such case the described differences, when applied to our California 

 material, do not differentiate two distinct units. Corolla length in relation to calyx is unstable in 

 this species group and often notoriously so in Polemoniaceae generally. Moreover, the presumed 

 pubescence character is slight and obviously variable in different collections otherwise alike. On 

 the basis of the known material such characters seem insufficient for use in separating two specific 

 units in the region of the South Coast Eanges ; and no other characters are available. 



Locs. — South Coast Eanges : Potrero HiUs, San Francisco (Zoe 2 :364) ; Woodside, San Mateo 

 Co., C. F. Baker 760; Mt. Hamilton, Eeller 7438; Loma Prieta, Santa Cruz Mts., Davy 371; Cold 

 Spr., Big Sur Eiver, Monterey Co., Jepson 2598 ; Tassajara Hot Sprs., Santa Lucia Mts., IF. I. 

 Follett 76; San Miguelito rancho, near Jolon, Monterey Co., Jepson 1642; La Panza, cent. San 

 Luis Obispo Co., Summers; Cuyama Valley, se. San Luis Obispo Co., Hoffmann. Western Mohave 

 Desert: Willow Sprs., Mnm 10,048. Cismontane S. Cal.: San Marcos Pass, Santa Ynez Mts., 

 Hoffmann (puberulent calyces and glabrous calyces occur on one individual) ; Agua Caliente, 

 Santa Ynez Mts., A. L. Grant 1681; Cow Caiion divide, San Gabriel Mts., Peirson 456; San Ber- 

 nardino Mts., Parish 462; Beaumont, Riverside Co., Jepson 6076; San Jacinto River near San 

 Jacinto, Jepson 1259; Hemet Valley, San Jacinto Mts., C. V. Meyer 765 (calyx glabrous) ; Tra- 

 buco Canon, Santa Ana Mts. (Abrams, Fl. Los Ang. ed. 2, 292) ; Lakeside, San Diego Co., K.Bran- 

 degee; Deseanso, San Diego Co., Wolf 2212. Great Valley, 10 to 500 feet (the usual plant is a 

 slight form, var. vallicola Jepson, with the following features : pedicels ascending, or often divari- 

 cate and curved at tip, the flower thus erect; calyx usually glabrous; corolla- throat commonly with 

 a puberulent or somewhat "frosty" zone at the stamen insertion near base of throat and below; 

 filaments usually hairy at base) : Willows, Glenn Co., Jepson 21,242 ; Main Prairie, e. Solano Co. 

 Jepson 20,935 (in both Main Prairie and Willows spms. the filaments are inserted in upper part 

 of corolla-throat, both the free and adherent parts glabrous) ; Byron, e. Contra Costa Co. C. F. 

 Baher 2864 ; Deadman Creek, n. of El Nido, Merced Co., Jepson 17,480 ; Shaf ter, Kern Co., Hoover 

 942; Famoso, Kern Co., Jepson 12,415 (individuals with glabrous calyces and individuals with 

 puberulent calyces occur in one narrow colony a few yards square, the plants plainly of one genetic 

 constitution) ; Lerdo, Kern Co., Jepson 17,343. Southern Sierra Nevada in Kern Co.: Greenhorn 

 Mts., Allison Krames. Tehachapi Mts.: Tehachapi, K. Brandegee. 



Refs. — Linanthus liniflorus Greene, Pitt. 2:254 (1892); Jepson, Fl. W Mid Cal 430 

 (1901), ed. 2, 334 (1911), Man. 803 (1925). Gilia liniflora Benth., Bot. Reg. sub t. 1622 (1833), 

 type from Cal., Douglas; DC, Prodr. 9 :315 (1845). G. liniflora subsp. eu-liniflora Brand ; Engler' 

 Pflzr. 4=™:133 (1907). G. pharnaceoides Benth., Bot. Reg. sub t. 1622 (1833), type from Cal.' 

 Douglas; DC, Prodr. 9:315 (1845). G. liniflora var. pharnaceoides Gray, Proc. Am. Acad. 8 :263 

 (1870). Linanthus pharnaceoides Greene, Pitt. 2:254 (1892) ; Jepson, Man. 803 (1925). Gilia 

 liniflora subsp. pharnaceoides Brand; Engler, Pflzr. 4"'':133 (1907). Var. vallicola Jepson 

 Man. 803 (1925), type loc. plain near Bakersfield, Jepson 8944. 



15. L. harknessii Greene. Stem strictly erect, very slender, 3 to 10 inches 

 high, branching mostly above the base and ending above in a dichotomously 

 branched paniculate cyme; herbage sparsely puberulent to glabrate; leaf -blades 

 palmately 3 to 5-parted into linear segments 1% to 8 lines long ; flowers minute, 

 solitary on filiform pedicels, the pedicels 3 to 10 lines long; ealj'x cupshaped or 

 turbinate, glabrous or nearly so, little shorter than the corolla, its lobes y^ to nearly 

 as long as the calyx-tube; corolla funnelform, white, % to 11/2 lines long, glabrous; 

 stamens inserted at the base of the corolla-throat, included in throat, glabrous; 

 capsule ovoid, the cells 1-seeded. 



Open places, usually in granitic soil, 3000 to 8000 feet : North Coast Ranges 

 from northern Lake Co. to western Siskiyou Co. ; Sierra Nevada from Fresno Co. 

 to Modoc and Siskiyou Cos. North to Oregon. June. 



Locs. — North Coast Eanges: Snow Mt., n. Lake Co., T. Brandegee; Soldier Ridge, Tollo Bolly 

 Mts., Jepson 20,932 ; Grouse Mt., Humboldt Co., Tracy 12,832 ; Big Flat, Salmon Mts., Hall 8644 ; 

 Shackelford Creek, w. Siskiyou Co., Butler 1657. Sierra Nevada: Jackass Mdw., South Fork San 

 Joaquin River, E. Ferguson 447 ; Huntington Lake, Fresno Co., Jepson 12,982 ; Fallen Leaf Lake 

 Eldorado Co., M. S. BaTcer ; Dog Valley, e. Nevada Co., Jepson 20,931 ; Summit sta., Nevada Co.' 

 Jepson 20,933; Webber Lake, Sierra Co., Lemmon; Jonesville, Butte Co., Heller 11,671; Meadow 

 Valley, Plumas Co., Jepson 10,646; Feather Lake, sw. Lassen Co., C. S. Sobinson; South Fork 



