246 HYDROPHYLLACEAE 



The type of P. magellanica has not been seen. On account, however, of the complex nature of 

 the problem and by reason of insufficient knowledge of the South American plants and of the type 

 of Phacelia magellanica, the name Phacelia magellanica, in view of these considerations, is reluc- 

 tantly adopted for the North American complex. The various North American forms occurring 

 in California are here described as follows : 



Var. calycosa (Gray) Jepson & Bailey comb. n. Stems stout, 1 to 2% feet high; herbage 

 hirsute; leaf-blades 1 to 6 inches long, pinnately divided, the lateral segments lanceolate, % to 

 IVi inches long, the terminal segment ovate to lanceolate, 1 to 3 inches long; petioles about % 

 to % as long as the blades ; racemes 2 to 4 inches long, borne on naked peduncles and disposed in 

 an open or sometimes dense panicle 3 to 14 inches long; corolla dingy white, campanulate, 2% to 

 4 lines long ; fruiting calyces ovate or broadly oblong, more or less imbricated, often conspicuously 

 so. — Dry rocky hills or flats, 100 to 3000 (or 4600) feet, mostly toward the interior; common 

 throughout cismontane California at low altitudes from Shasta Co. through the Sierra Nevada 

 foothills and Coast Ranges to San Diego Co. Apr.-June. 



Locs.- — Coast Ranges : Yreka, Butler 1356 ; Eighteen-mile Creek, Del Norte Co., Van Deventer 

 94 ; Horse Mt., ne. Humboldt Co., Tracy 7624 ; Van Duzen River Valley, Humboldt Co., Tracy 

 4112; Paskenta (5 mi. w.), sw. Tehama Co., Virginia Bailey; Stonyford, w. Colusa Co., Jepson 

 16,296; Willits, Mendocino Co., Docia Patchctt ; Round Valley, ne. Mendocino Co., Tracy 15,770; 

 Bartlett Mt., n. Lake Co., Jepson 18,922; St. Helena (e. of), Jepson 21,049; Dutton Canon, Vaca 

 Mts., Jepson 21,050; Pellejo Hills, w. of Vanden, Solano Co., Jepson 21,052 (intergrading to var. 

 heterophylla) ; Sonoma Valley, Jepson 4185 ; Benicia Hills, Jepson 18,884 ; Berkeley Hills, Jepson 

 9641; Mt. Diablo, Jepson 7611; Eva sta., e. slope Santa Cruz Mts., Jepson 21,054; Coyote sta., 

 Santa Clara Valley, Jepson 21,047 ; Little Arthur Creek, w. of Gilroy, Jepson 9706 ; Lorenzo Creek, 

 San Benito River, s. San Benito Co., Jepson 12,218; Mill Creek, Santa Lucia Mts., Jepson 1686; 

 Santa Margarita, San Luis Obispo Co., Summers. Great Valley: Redding, C. F. Base 277; Crane 

 Creek, w. Tehama Co., Jepson 21,056; Big Antelope Creek (2 mi. e. of Andrews road sta.), e. 

 Tehama Co., Jepson 16,627; College City, Colusa Co., Alice King; betw. Mossdale and Atlanta, 

 San Joaquin Co., Jepson 21,055. Sierra Nevada: Table Mt., Butte Co., Heller 10,786; Newcastle, 

 Placer Co., Bolander 4580 ; Sheep Ranch, Calaveras Co., T. Brandegee ; Table Mt., near Jamestown, 

 Tuolumne Co., Tracy 5696; Yosemite Falls, Yosemite, Jepson 4276 (calyx-lobes lanceolate); 

 Hogan Creek, Mariposa Co., Jepson 12,817; Switchback, Kaweah River, IV. Fry 337; Bear Creek, 

 s. Tulare Co., Furpus 1751; Democrat Sprs., lower Kern River, Jepson 19,896. Intramontane and 

 cismontane S. Cal. : Sisquoc River, n. Santa Barbara Co., If. S. Balcer ; Big Rock Creek, San Gabriel 

 Mts. (n. slope), Munz 6875; Eaton Canon, San Gabriel Mts., Peirson 164; San Bernardino, 

 Parish; Palomar Mt., T. Brandegee; Witch Creek, San Diego Co., Alderson 7570; Julian, Cuya- 

 maca Mts., Cleveland. 



Var. heterophylla (Pursh) Jepson comb. n. Stems stout, simple, 1 to 25 from a basal rosette, 

 usually erect, 1 to 3% feet high, densely leafy at base and leafy to the middle of the stem, thence 

 with the many shortly peduncled ascending racemes compactly virgate-racemose to the summit ; 

 stem and petioles sparsely or densely spreading-hispid, puberulent beneath; leaf -blades pinnately 

 divided to entire, appressed-pubescent ; calyx-lobes linear; corolla white, 2 lines long. — Dry hills, 

 rocky points and valley flats, (500 or) 2200 to 0900 feet: North Coast Ranges from Mendocino 

 Co. to western Siskiyou Co.; northern Sierra Nevada on the west slope from Mariposa Co. to 

 eastern Siskiyou Co., on the east slope from Mono Co. to Modoc Co. East to Nevada, north to 

 Oregon and Idaho. 



Geog. note. — The original specimen of Phacelia magellanica var. heterophylla (Pursh) Jep- 

 son, namely, Phacelia heterophylla Pursh, was collected by Meriwether Lewis "on irj hills on the 

 banks of the Kooskoosky" River, that is the present Clearwater River, opposite Kamiah, Idaho. 

 Of the original material, as preserved in the Herbarium of the Academy of Natural Sciences at 

 Philadelphia, only a portion of an inflorescence and a few leaves remain (Meehan, Proc. Acad. 

 Sci. Phila. 50:37). However, a collection, Constance, Hedrick & Peters 1897, made on Clearwater 

 River opposite Kamiah, is believed by Lincoln Constance to be the only Phacelia species growing 

 in the locality. If so, the number 1897 just cited is, therefore, to be regarded as a topotype. It is 

 this specimen, then, which furnishes the basis for the course here taken that Phacelia virgata 

 Greene of the California North Coast Ranges is in its fundamental characters quite the equivalent 

 of Phacelia heterophylla Pursh of Idaho, which species of Pursh is in this work treated as a 

 varietal subordinate of Phacelia magellanica. 



Of those forms of Phacelia magellanica which have a considerable distribution, var. hetero- 

 phylla (P. virgata Greene) is, in California, one of the most distinctive phases, though it inter- 

 grades with other forms on the margin of its distribution, especially with var. calycosa. Its 

 distinctive character is, however, emphasized by its capacity to preserve its main features not only 

 under a wide horizontal range but also under the climatic extremes of a wide altitudinal range. 

 As an example, it occurs at 500 feet in the Trinity River valley at Willow Creek (Tracy 13,794) 

 and, 19 miles distant, is found at 6900 feet on Salmon Summit (Tracy 14,385). These two locali- 

 ties fall within a narrow geographic area. 



Locs. — North Coast Ranges: betw. Burns Valley and Borax Lake, Lake Co., Hoover 3548; 

 Willits, cent. Mendocino Co., D. I. Patchctt ; Potter Valley, e. Mendocino Co., Purpus; Twin Rocks, 



