PHACELIA FAMILY 249 



type loc. Natoma ditch, Eldorado Co., K. Brandegee. P. magellanica f . ferruginea Brand ; Engler, 

 Pflzr. 4='>':100 (1913), type loc. Red Mt., n. Mendocino Co., Eastwood. Var. hetebophylla 

 Jepson. P. heterophylla Pursh, FI. 1:140 (1814), type loc. "on dry hilla on the banks of the 

 Kooskoosky," Ida., Lewis; Jepson, Man. 819 (1925). P. magellanica f, heterophylla Brand, Univ. 

 Cal. Publ. Bot. 4:218 (1912). P. virgata Greene, Erythea 4:54 (1896), type from "n. Cal. and 

 adjacent Ore.," that is, Yreka, Greene 832. P. virgata var. ampliata Greene, Pitt. 4:55 (1896), 

 type loc. Washoe Mts., Nev., perhaps belongs here. P. magellanica f. virgata Brand, Univ. Cal. 

 Publ. Bot. 4:219 (1912). P. calif omica var. virgata Jepson, Man. 820 (1925). P. calif ornica 

 t. vinctens Mcbr., Contrib. Gray Herb. 49:37 (1917), type loc. Donner Lake, Heller 6883. P. cali- 

 f ornica var. rubacea Jepson, Man. 820 (1925), type loc. Mt. Konocti, Lake Co., Jepson 21,063 

 (typ. in Herbario Jepsoniano). Var. californica Jepson & Bailey. P. calif ornica Cham., Lin- 

 naea 4:494 (1829), type loc. San Francisco, Chamisso ; Jepson, Fl. W. Mid. Cal. 438 (1901), ed. 

 2, 342 (1911), Man. 820, fig. 787 (1925). P. magellanica f. californica Brand, Univ. Cal. Publ. 

 Bot. 4:218 (1912). P. magellanica f. jepsonii Brand; Engler, Pflzr. 4==>i:100 (1913), type loc. 

 Mt. Tamalpais, Jepson 21,045. Var. qriseophylla Jepson. P. magellanica f . griseophylla Brand, 

 Univ. Cal. Publ. Bot. 4:218 (1912), tj-pe loc. Sisson, Siskiyou Co., K. Brandegee (typ. vidi). 

 P. heterophylla var. griseophylla Mcbr., Contrib. Gray Herb. 49:35 (1917); Jepson, Man. 819 

 (1925). P. heterophylla Jepson, I.e. ; not Pursh. P. mutabilis Greene, Erythea 4:55 (1896), type 

 loc. Castle Peak (first named locality), Nevada Co., Greene. P. nemoralis var. mutabilis Mcbr., 

 I.e. 49:37 (1917). P. stimulans Eastw., Proc. Cal. Acad. ser. 3, 2:291 (1902), type loc. Kings 

 Canon, Eastwood. P. magellanica t. egena Brand, Univ. Cal. Publ. Bot. 4:218 (1912), type loc. 

 Kaweah River, C. F. BaJcer i.i\5. P. californica f. egena Mcbr., Contrib. Gray Herb. 49:37 (1917). 

 P. californica var. egena Dundas, Bull. S. Cal. Acad. 33:161 (1935). P. nemoralis var. pseudo- 

 hispida Brand, Univ. Cal. Publ. Bot. 4:219 (1912), type loc. Stalker's, ne. Shasta Co., M. S. Baker 

 344, seems to be merely a form of var. griseophylla, probably produced by shaded habitat. P. pin- 

 nata var. pseudohispida Dundas, I.e. Var. cobymbosa Jepson. P. corymbosa Jepson, Man. 820 

 (1925), tjrpe loc. Dunsmuir, upper Sacramento River, Jepson 6161 (typ. in Herbario Jepsoniano). 

 Var. jAciNTENSis Jepson & Hoover. P. californica var. jacintensis Dundas, Bull. S. Cal. Acad. 

 33:160 (1935), type loc. Tahquitz Valley, San Jacinto Mts., Jaeger 1045. Var. patula Jepson. 

 P. magellanica f. patula Brand, Univ. Cal. Publ. Bot. 4:219 (1912), type loc. Stonewall Mine, 

 Cuyamaca Mts., Parish 4423. P. californica var. patula Jepson, Man. 820 (1925). Var. fkigida 

 Jepson. P. frigida Greene, Pitt. 4:39 (1899), type loc. Mt. Shasta, C. H. Merriam. P. alpina 

 Rydb., Mem. N. Y. Bot. Gard. 1:324 (1900), type loc. Cedar Mt., Mont., Eydberg 4- Bessey 4855. 

 P. magellanica f. alpina Brand, Univ. Cal. Publ. Bot. 4:217 (1912). P. magellanica f. frigida 

 Brand, I.e. 218. P. heterophylla f. frigida Mcbr., Contrib. Gray Herb. 49:35 (1917). P. hetero- 

 phylla var. frigida Jepson, Man. 819 (1925). P. leucophylla var. alpina Mcbr. I.e. 34, as to Cal. 

 plant cited. P. compacta Greene; Baker, West Am. PI. 1:18 (1902), type C. F. Baker 1142 

 [Spooner, Douglas Co., Nov.] ; nomen nudum. P. magellanica f. compacta Brand, Univ. Cal. 

 Publ. Bot. 4:217 (1912). P. leucophylla var. compacta Mcbr., Contrib. Gray Herb. 49:34 (1917). 

 P. heterophylla var. dasyphylla Jepson, Man. 819 (1925), type loc. Silver Pass, Fresno Co., A. L. 

 Grant 1532 (first adequate publication), the name based on P. dasyphylla Greene (Brand in Eng- 

 ler, Pflzr. 4-''' :97, — 1913, nomen subnudum) ; the filaments vary from slightly hairy to glabrous in 

 high alpine material from the Mt. Whitney region ; since there are no other differences between 

 this form and var. frigida, such a slight variation is negligible. P. heterophylla var. pygmaea 

 Jepson, Man. 819 (1925), type loc. Mt. Shasta, Jepson 20,992 (typ. in Herbario Jepsoniano). 



3. P. nemoralis Greene. Stems 1 or few from the base, simple below, panicu- 

 lately branched above, I14 to 3 feet high, very bristly with long stinging hairs ; 

 herbage light green ; leaf -blades elliptical to oblong, 1 to 4 inches long, simple and 

 entire or with a pair of small leaflets at base ; basal and lower leaves on petioles 

 2 to 3 inches long, uppermost short-petioled or sessile; racemes IV2 to 2i/2 inches 

 long, slender, in 2s or 3s, terminating the stems or lateral branches ; corolla whitish, 

 2 lines long; capsule 2-seeded. 



Moist shady flats or hillslopes, 20 to 1800 feet : Coast Ranges from the Santa 

 Cruz Mts. to Mt. Diablo and Sonoma Co. June. 



Tax. note. — Phacelia nemoralis differs only slightly from various forms of Phaeelia magel- 

 lanica and on the basis of morphological characters were perhaps better disposed as a varietal 

 form of that complex or reduced to var. griseophylla of P. magellanica. The narrower-leaved 

 states of Phacelia nemoralis (Mt. Tamalpais, Eastwood; Oakland Hills, Lemmon; Loma Prieta, 

 W. W. Price) simulate closely Phacelia magellanica var. griseophylla. It must be noted, too, that 

 of the varieties of Phacelia magellanica, var. frigida, for example, differs more from var. calycosa 

 (the most abundant and geographically central phase in California) than does Phacelia nemoralis 

 from the var. calycosa. This circumstance is, to be sure, often a feature of complex species, a 

 great diversity within the group accomp.anied by a wide mass of intergradation. Phacelia nemo- 

 ralis is, however, biennial as opposed to the perennial duration of all Phaeelia magellanica forms 

 and if it be narrowly restricted to the phase represented by Greene's original specimens, then it 



