BORAGE FAMILY 363 



Allocarya californica has been described as having the nigae on the ventral side of nutlets distinct 

 from keel as contrasted with the rugae on ventral side joined to the keel or closely approaching 

 it in A. eusickii var. vallicola. This distinction has been proposed as a differentia but it is difficult 

 in most cases to apply and is not thought significant. Moreover, for A. californica there are 

 obscurities in another direction. Differentiae between A. californica (especially as represented 

 by Humboldt County plants) and the Alaskan A. picbeia Greene (Lithospermum plebeium Cham., 

 Linnaea 4:446, — 1829, type loc. Uualaska, Alas., Chamisso) have not as yet been clearly estab- 

 lished. It seems wiser, however, at this stage of investigation, to retain the concept Allocarya 

 californica aa here set up for this group of the California coast line, a group which is fairly defi- 

 nite in its main features. 



In Humboldt Co., the ordinary state of A. californica passes into a strain, var. retictjlata 

 (Piper) Jepson comb, n., in which the nutlets are delicately circular-reticulate or pentagonal- 

 reticulate or quadratish-reticulate, w^th the areolae smooth or sometimes finely papillate: Bear 

 Eiver (10 mi. from ocean), Tracy 15,351; Eureka, Tracy 2111; Butler Valley, Mad River, Tracy 

 5825; Bald Mt., Tracy 15,009; Big Lagoon, Tracy 13,923. Since the intergrades are many, this 

 form is scarcely worth separation or notation. The next variety is more important since the 

 character of the nutlet is associated with a very slender spike and a large corolla. 



Var. mlnuta (Piper) Jepson & Hoover comb. n. Stems 1 or several from the base, simple or 

 branching, erect, slender, 6 to 10 inches high; calyx small (% to 1 line long) ; corolla large in 

 proportion to the calyx (2 lines broad); nutlets small (% to % line long), irregularly tine- 

 reticulate over the entire back or only at the middle, not tuberculate. — Wet places in valleys, 350 

 to 1000 feet : Eel River near Fort Seward. In its nutlets this form is very close to var. reticulata. 



Refs. — Allocarya CALiroRNiCA Greene, Pitt. 1:20 (1887), as to the name, which rests on 

 Myosotis californica F. & M., Ind. Sem. Hort. Petrop. 2:42 (1835), a transcript of which is 

 printed in Erythea 2 : 149, — 1894. The type locality for Myosotis californica is given as Fort Ross, 

 Sonoma Co. It is true, in the case of the Russian colony, that some of the plants of the Russian 

 collectors were gathered on expeditions into the interior — but failing contrary evidence, as in 

 this instance, the specific locality Fort Ross must be assumed as authentic, since the plant here 

 believed to be Myosotis californica F. & M. grows near Fort Ross. Gray transferred Myosotis 

 californica to Krynitzkia but applied the binomial to the inland species mainly. Greene, too, had 

 chiefly in mind the interior plant. In publishing Allocarya californica, he cited the original pub- 

 lication of Myosotis californica F. & M. and indicated as range, "from the coast to the foothills of 

 the Sierra Nevada," thus, in theory at least, including the coastal plant. His binomial must, 

 therefore, nomenelatorially, go to the coastal species, although actually, in practice, he applied 

 the name mainly and probably exclusively to the plant of the interior, here taken to be a dif- 

 ferent species. 



The synonymy of Allocarya californica (F. & M.) is as follows: Eritrichium califomicum 

 A.DC, Prodr. 10:130 (1846). Krynitzkia californica Gray, Proc. Am. Acad. 20:266 (1885), in 

 part. A. diffusa Greene, Pitt. 1:14 (1887), type loc. U. S. Marine Hospital, San Francisco, 

 Greene; Jepson, Fl. W. Mid. Cal. 443 (1901), Man. 853 (1925). Flagiobothrys diffus-us Jtn., 

 Contrib. Arn. Arb. 3:77 (1932). A. scalpta Piper, Contrib. U. S. Nat. Herb. 22:104 (1920), type 

 loc. Alder Point Flat, Humboldt Co., Tracy 1878; Jepson, Man. 852 (1925). A. areolata Piper, 

 I.e. 22:105 (1920), type loc. Mendocino, Pringle. A. dispar Piper, I.e. 22:109 (1920), type loc. 

 Agnes, Curry Co., Ore., /. C. Nelson 1433. Flagiobothrys reticulai us var. rossianorum Jtn., 

 Contrib. Arn. Arb. 3:79 (1932), name based on Myosotis californica F. & M. Var. reticulata 

 Jepson. A. reticulata Piper, I.e. 22 : 105 (1920), type loc. Holmes Flat, Eel River, Humboldt Co., 

 Trac)/ 4691. ^. mtTiiita var. reitcu^aia Jepson, Man. 852 (1925). Flagiobothrys reticulatus Jtn., 

 Contrib. Arn. Arb. 3:78 (1932). Var. minuta Jepson & Hoover. A. minuta Piper, Contrib. 

 U. S. Nat. Herb. 22 :104 (1920), type loc. Fort Seward, Humboldt Co., Tracy 4469; Jepson, Man. 

 852 (1925). 



11. A. eusickii Greene. Stems several to many from the base, erect or diffuse 

 or decumbent, 4 to 10 inches high; herbage ascending-hispid or spreading-hispid; 

 leaf -blades linear, I/2 to IV2 inches long ; spikes i/4 to 1 inch long; calyx only slightly 

 accrescent ; corolla Y2 to 1 V2 lines broad ; nutlets ovate, a little contracted or attenu- 

 ate above the middle to a short lanceolate beak, slightly flattish dorsally and ven- 

 trally with the beak a little obeompressed, transversely rugose dorsally and par- 

 tially areolate, the intervals minutely papillate; ventral side keeled; scar linear, 

 strictly lateral but reaching base of nutlet, usually set in a broad depression. 



Montane flats and meadows, 4000 to 6100 feet : easterly summits or east side of 

 the Sierra Nevada from Placer Co. to Modoc Co. East to Utah and north to Oregon. 

 June. 



Locs. — Summit Soda Sprs., Placer Co., ShocMey 322 ; betw. Vinton and Beckwith, e. Plumas 

 Co.; Honey Lake, T. Brandegee ; Goose Lake Valley, Modoc Co., K. M. Austin; Twelve-mile Creek, 

 Warner Mts., L. S. Smith 1259. 



