426 



LABIATAE 



kind), continuous variability in leaf shape and size (size being often associated with ecological 

 situation). There is no constant correlation of the variables, pubescence and leafage, but, on the 

 other hand, the entire assemblage is characterized by essentially like habit, by essentially like 

 spikes, by like flowers and by like nutlets and is therefore evaluated as one species by reason of the 

 likeness in constant characters that must be regarded as important. In nature, as to this group, 

 pubescence and leaf shape and size are not only expressed by variables but the variables may be 

 partially developed into certain extremes. An attempt is here made to segregate the pubescence 

 and leaf extremes of this aggregate under varietal names. The citation of stations under such 

 varietal names is, however, often attended with such dilemmas as to compel resort to a decision on 

 geographic grounds rather than on morphological grounds — a concession to the complexity of the 

 web of variables. Such proposed varietal units, as those listed below, are, in 

 consequence, somewhat artificial. 



The following collections represent intergrades from the species to the 

 variety quercetorum : Ross Valley, Marin Co., Jepson 14,957 ; La Jota plateau, 

 Howell Mt., Napa Range, Jepson 17,575; Fort Bragg, Mathews. There is, 

 therefore, next described : 



Var. quercetorum (Hel.) Jepson. (Fig. 439.) Herbage green, hir- 

 sute or hirsutulose (mostly sparsely so), frequently also puberulent and 

 often glandular, the stem angles hirsute or often retrorsely hirsute or short- 

 hispid ; leaf -blades broadly to narrowly ovate (at least the lowermost), 1 to 

 2 (or 3) inches long, cordate to truncatish at base, more sparsely hairy than 

 in the species, the margins with larger teeth; petioles of the lower leaves 

 mostly Va to 1% (or 2) inches long. — Openly wooded dry hUls, moist valley 

 flats and canon bottoms, 10 to 2500 feet: coastal Southern California from 

 San Diego to San Jacinto Valley and Santa Monica; Coast Ranges from 

 Santa Cruz Co. to Humboldt Co. South to northern Lower California. The 

 commonest form in the central Coast Ranges, but largely or wholly absent 

 over large areas from Monterey Co. to Orange Co., where it is replaced by 

 Stachys buUata. 



Locs. — S. Cal. : La JoUa, Jepson 11,884; Claymine Caiion, Santa Ana 

 Mts., Howell 2644; San Jacinto, Basse; San Bernardino, Parish; Griffith 

 Park, Los Angeles, Braunton 544; Santa Monica, Basse. Coast Ranges: Los 

 Gatos (hills w.), Beller 7214; Stanford, Airavis 2366; Los Buellis Hills, 

 Mt. Hamilton Range, B. J. Smith; Livermore Valley, Jepson 14,954; Berke- 

 ley, Jepson 14,983; Button Caiion, Vaca Mts., Jepson 14,980; St. Helena 

 (hills e.), Jepson 14,982; Kelseyville, Lake Co., Jepson 14,978; Fort Bragg, 

 W. C. Matheivs; Dyerville, mouth of South Fork Eel River, Jepson 16,684; 

 Kneeland Prairie, Humboldt Co., Tracy 3881. 



There are many intergrades from var. quercetorum to the next variety, 

 var. rigida. There are also intergrades from the species to var. rigida (such 

 as Patricks Creek, Del Norte Co., Paries # Tracy 11,405). 



Var. rigida (Nutt.) Jepson & Hoover comb. n. Stems erect, mostly 

 simple, hirsute with spreading hairs, 1 to 3 feet high ; leaf -blades ovate or 

 oblong-ovate, serrulate or crenulate, mostly rounded or truncatish at base, 

 acute or acuminate at apex or sometimes obtuse, hirsute or silky, 1 to 3^/4 inches long; petioles % 

 to 1% inches long. — Stream banks and bottoms, moist flats and meadows, 3500 to 6500 feet: 

 mountains of Southern California; northern Sierra Nevada from Mariposa Co. to Modoc and 

 eastern Siskiyou Cos. ; Coast Ranges from Humboldt Co. to western Siskiyou Co. North to Oregon. 

 June-July. 



Locs. — S. Cal.: Noble Mine, Laguna Mts., Chandler 5497; Cuyamaca Mts., T. Brandegee ; 

 Saunders Mdws., San Jacinto Mts., C. V. Meyer 585 ; Strawberry Valley, San Jacinto Mts., Jepson 

 2270 ; San Bernardino Mts., Parish 1725. Northern Sierra Nevada : Yosemite, Ball 9225 ; Douglas 

 Flat, e. Tuolumne Co., A. L. Grant 103; Morgan Sprs., Tehama Co., Jepson 12,325; Goose Lake, 

 Modoc Co., R. M. Austin. In the foregoing the herbage has, commonly, a somewhat yellowish or 

 brownish cast, the leaf -blades oblong-ovate and mostly obtuse or obtusish. In Lassen and Tehama 

 Cos., the following are less hairy : Martin Sprs., near Eagle Lake, Brown f Wieslander 57 ; Mineral, 

 J. Grinnell. In Humboldt, Trinity, Siskiyou and Del Norte Cos., the herbage is usually green, 

 often very thinly hirsute, the leaf-blades commonly acute but sometimes obtuse, the spikes fre- 

 quently much elongated: Grouse Mt., Humboldt Co., Tracy 13,455 ; Snow Camp Mt., Humboldt Co., 

 Tracy 15,652; Trinity Summit, Tracy 15,091; Shasta Sprs., Siskiyou Co., Jepson 14,958; Woolley 

 Creek, Marble Mt. region, Butler 172; Yreka, Butler 173; Dunn Creek, ne. Del Norte Co., Jepson 

 18,515. 



In the form called Stachys rivularis Hel., the herbage is green, thmly hirsute, scarcely ever 

 silky ; the leaves mostly narrowly oblong, sometimes oblong-ovate, serrulate, 1% to SVa inches long, 

 shortly petioled below, subsessile or sessile above the base. — It is found on the margins of marshes 

 or along streams, 3000 to 6500 feet, in the northern Sierra Nevada from Tuolumne Co. to eastern 

 Siskiyou Co. and Modoc Co. It is a narrow leaved or oblong-leaved form of var. rigida in which 



Fig. 439. Sta- 

 chys AJUQOIDES 

 Benth. var. quer- 

 cetorum Jepson. 

 fl. branchlet, X 1. 



