MINT FAMILY 



435 





*^^^±i:s. 



as far as Mariposa County. Within this distribution area it has differentiated a considerable num- 

 ber of forms. Though these forms have not developed distinctive structural characteristics suffi- 

 cient to mark them as separate entities, a certain portion or phase of each form that is most strongly 

 or evidently differentiated is, for convenience, here described under a varietal name. While these 

 varieties, so-called, are segregated geographically — some to a marked degree, some scarcely at all — 

 it is to be emphasized that blocks or areas of intermediates or of shadowy divergences here and 

 there tend to disqualify the described varieties which are in reality uneasy variants, and in this 

 sense they are merely nominal. It is significant to observe that the degree of differentiation in 

 the Coast Ranges, with its marked microclimatic areas, is very much greater than under the more 

 uniform conditions of the Sierra Nevada footliills, where the representation is almost confined to 

 a single slight but fairly constant phase, the var. glabella. 



This species, Monardella \'illosa, was discovered at Bodega, Sonoma County, by E. B. Hinds 

 surgeon and botanist of the British exploring ship Sulphur, in 1837. His collection is preserved at 

 Kew. The leaves in the type are not truncatish or sub- 

 cordate at base as in the littoral var. franciscana but ob- 

 tuse; they are hairy-pubescent beneath but not felt-like 

 or white-tomentose as in var. franciscana (cf. Jepson, 

 Calif ornian Types at Kew, 294-295. 1906. ms.) The type, 

 therefore, is represented by plants which lie, for the most 

 part, back of the coast line or well towards the interior 

 and are not strictly littoral. This representation, the 

 typical form, is to all appearances the most widely dis- 

 tributed manifestation of the aggregate. Of it citations 

 are set down as follows. 



Locs. — San Carpojo, San Luis Obispo Co., Gondii; 

 Lorenzo Creek, upper San Benito River, Bettys; Carmel 

 River, Parish 20,043 ; Glen Echo, Santa Cruz coast, Jep- 

 son 59g; Arroyo Mocho, n. end Mt. Hamilton Range, 

 Jepson 14,986 ; Morrison Caiion, Niles, Jepson 59h ; Mt. 

 Diablo, Bower man 339 ; Skyline Ridge, Berkeley Hills, 

 Jepson 18,437; betw. Guerneville and Monte Rio, Russian 

 River, E. Ferguson 221 ; Fort Bragg, Mathews 169; Gar- 

 bervDle, s. Humboldt Co., Jepson 12,370 ; Grasshopper 

 Peak, head of Canoe Creek, South Fork Eel River, Jepson 

 16,483 ; Dyerville, mouth of South Fork Eel River, Jepson 

 16,433 ; Somes Bar, Klamath River, Chandler 1522. 



Var. franciscana (Elmer) Jepson. Stems reclining 

 or diffuse, more or less woolly -pubescent; leaves broadly 

 ovate, serrate to entire, pubescent above, especially the 

 upper ones, white-woolly beneath, % to 1% inches long. — 

 Coast line from Pt. Reyes to San Luis Obispo, 5 to 100 

 feet: Alamere Creek, 8 mi. n. of Bolinas, Merle Sandall; 

 betw. Mussel Rock and Salada, San Mateo Co., Neulon 

 253; Pajaro Hills, Chandler 366; San Luis Obispo (coast 

 hills near), Snmmers. June-July. 



Var. obispoensis Hoover var. n. Leaves round-ovate, usually entire, sometimes crenate, 

 densely white tomentose below, in strong contrast to the pubescent but green upper surface, hairs 

 all branched ; corolla white. (Folia ovata, plerumque integra, infra dense albo-tomentosa, supra 

 viridia ; capilli ramosi.) — Western San Luis Obispo Co.: Cuesta Pass (near). Gondii (type); 

 Perfumo Caiion, Unanpst; Pine Mt., near San Simeon Bay, Palmer. The branched hairs are a 

 remarkable feature. They have been observed in no other Monardella. A plant of the Nacimicnto 

 River, sw. Monterey Co., Peirson 11,829, grades towards this variety. 



Var. globosa (Greene) Jepson comb. n. Leaf-blades broadly ovate, serrate, coarse, venose, 

 thinly pubescent or tomentulose beneath when young, 1 to 1% inches long; heads large (1 to 1% 

 inches broad), the outer bracts foliaceous, conspicuous, strongly reflexed. — Hills on east side of 

 San Francisco Bay, 50 to 800 feet : Oakland Hills, Drew; North Berkeley Hills, E. A. Walker 605 ; 

 Martinez, K. Brandegee. 



Var. tomentosa (Eastw.) Jepson. Stems and lower surface of leaves (especially the upper 

 ones) and bracts, also the calyx, covered with soft gray pubescence ; bracts foliaceous; heads ter- 

 minal or sometimes smaller ones in the upper axils. — Dry flats or hillslopes from the Napa Range 

 to central Mendocino Go. : Calistoga, Jepson 9969 ; Laytonville. 



Var. neglecta (Greene) Jepson comb. n. Stems 4 to 14 inches high; herbage glabrous or 

 microscopically puberulent, the stems, leaves and heads often purplish ; leaf-bladc^s ovate, serrulate, 

 dark green, purple-veined, those of the basal sterile shoots 2 to 5 lines long, those of the flowering 

 stems often larger (4 to 7 lines long) ; heads ijiclincd to be small (6 to 12 lines broad) ; corolla 

 purple. — Rocky slopes, Marin and Sonoma Cos.: Tiburon, Jepson 12,920; Mt. Tamalpais (Wheeler 

 trail, Howell 5382; Rock Spring trail, K. Brandegee) ; betw. Graton and Occidental, Hoover 3609. 



Fig. 441. Monardella villosa 

 Benth. a, fl. stem, X 1 ; b, fl., X 1% ; 

 c, calyx X %. 



