452 



SOLANACEAE 



(monochasial cymes), the bracts (except the lower) mostly small; calyx cleft 

 nearly half way into lanceolate lobes, about half as long as the corolla; corolla 

 greenish-white or greenish-yellow, 8 to 11 lines long, constricted just beneath the 

 spreading limb, the limb at first slightly cupped or finally somewhat reflexed, 31/2 

 to 6 lines broad ; filaments hairy at very base, inserted at summit of corolla-tube. 

 Canon walls, rocky outcrops on mesas and in washes, 5 to 4000 feet : Mono and 

 Inyo Cos. ; Mohave and Colorado deserts. East to Texas, south to Mexico. Apr. 



Note on duration. — The stems arise eacli season from the marlcedly woody root-crown of a 

 perennial taproot. Often the slightly sxiffrutescent bases of the stems live over a few winters. 



In all the deserts this author has found Nicotiana trigo- 

 nophylla perennial. However, in the botanic garden at 

 Berkeley it is, ace. Helen-Mar Wheeler, generally annual 

 under cultivation. Over most of Arizona it is, too, an- 

 nual but sometimes perennial in the southwestern part 

 of the state (ace. Kearney & Peebles). 



Locs. — Mono Co.: Painted Caiion, Curtis Ferris 67. 

 Inyo Co.: Texas Spr., Funeral Mts., Jepson 6884; John- 

 son Caiion, Panamint Range, Jepson 19,625. Mohave 

 Desert: Granite Mts. (near Avawatz Mts.), Jepson 

 17,285; Paradise Mts. (41/2 mi. ne.), Jepson 20,334; 

 Willow Spr. (cove below). Old Dad Mts., Jepson 20,424; 

 Ord Mt., Jepson 5853; Needles (10 mi. w.), Newlon 543. 

 Colorado Desert: Chuckwalla Wash, e. Riverside Co., 

 Schellenger 91 ; Painted Canon, Mecca Hills, Jepson 

 11,651; Palm Canon of Mt. San Jacinto, Jepson 1409; 

 San Felipe Wash, e. San Diego Co., C. V. Meyer 75; 

 Vallecito, e. San Diego Co., Jepson 8627; Mountain 

 Springs grade, w. Imperial Co., Newlon 375; Milpitas, 

 Colorado River, Jepson 5284. 



Most of the California collections of Nicotiana tri- 

 gonophylla approach N. palmeri Gray (Syn. Fl. 2:242,— 

 1878, type loe. Williams Fork, n. Ariz., Palmer 433) 

 which is reduced to N. trigonophylla var. palmeri by 

 Jones (Contrib. W. Bot. 12:52,— -1908). Plants of typi- 

 cal N. palmeri are more robust than N. trigonophylla, 

 appear to be of slower growth and lower, the inflorescence 

 tends to be shorter and more leafy, whUe the corolla is 

 not only larger but whiter, the limb well-spread and dis- 

 tinctly lobed. — Helen-Mar Wheeler. 



Refs. — Nicotiana trigonophylla Dunal in DC, 

 Prodr. 13: 562 (1852), type loc. Aguas Calientes, Mex., 

 Hartioeg; Jepson, Man. 888 (1925). 



3. N. acuminata Hook. var. multiflora Reiche. (Fig. 446.) Stem erect, 

 branching, rather coarser than in N. attenuata, commonly I14 to 5 feet high; 

 herbage strongly glandular-pubescent, ill-smelling; leaf -blades ovate to narrowly 

 ovate-lanceolate, acute or acuminate, 2 to 10 inches long, obtuse, truncatish or sub- 

 cordate at base; branches of the panicle elongated, spreading, loosely racemose; 

 calyx-tube with 5 narrow longitudinal dark bands opposite to and extended up the 

 lobes; calyx-lobes linear or subfiliform, equaling or exceeding the tube ; corolla 1% 

 to 2 inches long, the throat tubular with spreading limb 6 to 10 lines wide, the tube 

 proper strongly narrowed below the calyx-teeth ; stamens inserted at summit of 

 the corolla-tube proper. 



Introduced from South America and naturalized in valley flats and creek "bot- 

 toms," 50 to 500 (or 4500) feet : scattered stations in central and northern cismon- 

 tane California. June-Oet. 



This plant, Nicotiana acuminata var. multiflora, has been spontaneous as an alien immigrant 

 in California for full fifty years, probably sixty years, perhaps longer. It has been misdeter- 

 mined by botanists (including the author) as Nicotiana attenuata which it superficially resembles. 

 The territory which it occupies in central California is quite distinct from the territory inhabited 

 by Nicotiana attenuata. Investigating the genus Nicotiana grnerally and correcting a previous 

 misdetermination of this species, W. A. SetcheU was without any doubt the first botanist to re- 



Pig. 446. Nicotiana acuminata 

 Hook. var. multiflora Reiche. a, fl. 

 branchlet, X % ; 6, leaf, X % ; c, 

 long. sect, of corolla-tube and -throat, 

 X 1. 



