26 CRUCIFERAE 



Locs. — Bakersficld, Greene; Delano, Otto Steinwand; Deer Creek, Tulare Co., Congdon; 

 Tulare (Fl. Fr. 25C) ; Huron, Fresno Co., T. Brandegcc ; Alcalde Canon (n. side) ; Zapato Chino 

 Creek, T. Brandcgce (Zapalo Oliiuo Creek is tlie full name of Zapato Creek, a winter water course 

 on the east slope of the Diablo Range, sw. Fresno Co.) ; Lost Hills (5 mi. e.). Mason; Carrizo 

 Plain, opp. Panorama Hills, se. San Luis Obispo Co., Jcpson 10,214. 



Eefs. — Streptanthu.s cai.ifoknious (ireene, Fl. Fr. 250 (1891) ; Jepson, Man. 417 (1925). 

 Stanfordia calif ornica Wats. Bot. Cal. 2:479 (1880), type loc. near Tulare, Mrs. A. E. Bush. 

 Caulanthus calif orniciis Payson, Ann. Mo. Bot. Card. 9:299 (1922). 



9. S. insignis Jepson. Annual; stem branching from near the base, 1/3 to l^/^ 

 feet high; herbage bristly-pilose; raceme with a dense terminal cluster of sterile 

 flowers forming a conspicuous black spot; pedicels bristly-hispid; calyx sparsely 

 but markedly bristly-hispid; stigma evidently 2-lobed; pods bristly-hispid, erect 

 or reflexed. 



Stony mountain slopes, 1500 to 2500 feet; inner South Coast Range from San 

 Benito Co. to w. Fresno Co. Apr.-May. 



Tax. note. — While the stigma of Strcptanthus insignis is only slightly 2-lobed, yet the loba- 

 tion is obvious. This species is probably a mutant from S. coulteri Greene. It is, in any event, 

 on the one hand, by reason of its hairy pedicels, the terminal "black spot" on the raceme, and its 

 stigma, closely related to S. coulteri; and we believe that it is, on the other hand, related to 

 S. glandulosus var. pulchellus Jepson by reason of its flowers. We find that collectors have dis- 

 tributed specimens of S. insignis under the name S. pulchellus Greene, but that plant has a cir- 

 cular and quite entire stigma. The genetic position of Strcptanthus insignis is, indeed, strikingly 

 intermediate between S. coulteri Greene and S. glandulosus Hook., though nearer the former, and 

 gives further weight to the soundness of the position we here hold that these two species, Strcptan- 

 thus glandulosus Hook, and Strcptanthus coulteri Greene (Caulanthus coulteri Wats.), do not 

 belong to different generic units. 



Locs. — Se. San Benito Co., Hall 9941; Warthan, \v. Fresno Co., Eastwood; Pinnacles, Mon- 

 terey Co., A. J. Pietcrs (some of the pods glabrous, the filaments distinct). 



Ref. — Streptajstthus insignis Jepson, Man. 420 (1925), type loc. Warthan, w. Fresno Co., 

 Eastwood. 



10. S. coulteri Greene. Annual; stem simple or branching, 1 to 3 feet high; 

 herbage thinly hispid; blades of cauline leaves oblong to ovate or lanceolate, entire 

 or dentate to pinnately lobed or parted, sagittate-clasping, 1 to 5 inches long, 

 blades of the basal and lowest cauline similar, on short winged petioles; flower-buds 

 congested, deep purple, showing as a terminal black-purple spot at the summit of 

 the plant, the racemes therefore at first very dense, but soon becoming elongated 

 and lax; flowers 6 to 7 lines long, spreading or pendulous, on pedicels 1 to 5I/2 lines 

 long; pedicels and calj'x more or less bristly; calyx at first deep purple, becoming 

 greenish; petals white, dark veined, widely spreading; filaments of longest pair 

 of stamens united about % or % of their length, or sometimes all distinct; pods 

 terete, stoutish, erect, reflexed or reflexed-spreading, glabrous, 2^/2 to 3 inches long. 



Interior foothills and valley floors, 250 to 4000 feet : upper San Joaquin Valley 

 and neighboring foothill ranges and foothill valleys, southward to the mountains 

 on the west side of the Mohave Desert. Mar.-May. 



Distribution note. — Strcptanthus coulteri occurs chiefly in small colonies with well defined 

 boundaries and to a less degree as scattered individuals. The known colonies are comparatively 

 few in number. In 1932 a colony was found in Palo Prieto Pass, inner South Coast Eange, on 

 a westerly slope, the plants (in full flower) standing so thickly as to make a light-colored spot 

 on the hill about 80 by 100 feet in area. All the individuals were plainly of one close descent 

 as evidenced by the uniformity of habit, aspect, leaves, inflorescence and flowers. The flowering 

 axes in the course of development of the fruit showed successively pendulous, spreading and 

 finally erect pods. The half -grown pods are curiously vermiform-contorted. 



Note on the flower. — The sepals are strongly keeled. The broad claws of the petals are 

 contracted at apex and expand into the very undulate limbs. The limbs are really broader than 

 the claw, but, being troughed and strongly undulate, do not appear so ; they are white, or nearly 

 so,..with a few very dark or black pinnate veins. The short stamens are only half the length of 

 the long stamens. The pedicels are hirsute with spreading hairs ; there are also a few hairs of 

 this kind advanced on the calyx keels. The pedicels and calyces of the congested flower-buds at 



