MUSTARD FAMILY 27 



the tip of the raceme are glabrous ; these upper flowers with glabrous buds are sterile ; the calyces 

 of the older flowers are sparsely hirsute or bristly. Pubescence, however, varies in amount and 

 may be absent from individual organs. 



The embryos are very remarkable. Both cotyledons are 

 deeply parted into three rounded lobes as shown in fig. 130. 

 These remarkable structures afford further evidence of great 

 weight that Streptanthus coulteri and S. californicus are 

 closely allied. See also the discussion on the embryo in the 

 taxonomic note under Streptanthus californicus. l '^ 



Locs. — Sierra Nevada foothills from Madera Co. to Kern (7 t/ O 



Co.: Pollasky, "Madera Co." (Ann. Mo. Bot. Gard. 9:297) ; Fig. 130. Streptanthus COTIL- 



Dunlap, Fresno Co., Jepson 2760; Orosi, Tulare Co., Harriet teri Greene, a, b, embryos, X 6; 

 Kelley; Kaweah, Hopping 270 ; Havilah, Bear Creek, Purpus c, cotyledon spread out, X 6. 

 1705 ; Caliente, Davy 1880. San Joaquin plain : Tulare, Davy; 



Oil City, Kern Co., Heller 7630 (pods erect). Inner South Coast Range: Palo Prieto Pass, e. 

 San Luis Obispo Co., Jepson 16,201; White Hills, Cuyama, Eastwood. Mt. Pinos region: Bisses 

 sta., Tehachapi Mts., Dudley 455 (pods spreading) ; San Emigdio, Eastwood; Leonis Valley 

 Davy 2634; Saugus, Davy. 



Var. lemmonii Jepson comb. n. Habit of the species; herbage glabrous or sparsely hirsute 

 on lower part of stem; leaf -blades entire, coarsely toothed, or only weakly denticulate; raceme 

 rather loose, the flowers deflexed or pendulous on pedicels 2 to 4 lines long ; pedicels glabrous or 

 the lower ones hairy; calyces glabrous or the lower ones hairy; pods erect, stout, 2 to 4 inches 

 long; stigmas long, spreading conspicuously. — Dry rocky banks, 900 to 1200 feet: San Luis 

 Obispo Co. Apr. 



Locs. — Paso Eobles, Barter, Gondii; Cholame, Lemmon. This form has not been re-collected 

 in the Cholame region. This fact in connection with consideration of the climatic differences 

 between the Santa Lucia Mts. and the Inner South Coast Range suggests the surmise that Lemmon 

 collected his plant near Paso Robles while on one of his trips from Paso Robles to Cholame. We 

 think it likely, therefore, that var. lemmonii is a very narrow endemic restricted to the Paso Robles 

 region. 



The flowers are soon pendulous, likewise the very young fruits, which, however, become erect 

 in maturity. In general only the lower flowers are fertile. In all of this there is close resemblance 

 to S. coulteri. The distinguishing marks of S. lemmonii Wats, were supposed to be the glabrous 

 herbage, the glabrous pedicels and calyces, the deeply cleft stigma and the erect pods. But erect 

 pods may occur in S. coulteri and as to pubescence, the following forms show the range of vari- 

 ation in hairiness: (1) The extreme form, herbage, pedicels and calyx quite glabrous save that 

 the stem is slightly hispid at base (Paso Robles, Barber). (2) Lower pedicels hairy, calyces 

 glabrous, stems hirsute at base (Paso Robles, comm. Georgiana P. Ballard). (3) Lower pedicels 

 densely bristly; calyces of lower flowers hairy; stem hirsute below (Paso Robles, comm. Geor- 

 giana P. Ballard). (4) All the pedicels hairy except the purple ones of the sterile cluster; calyces 

 mostly glabrous (Paso Robles, Carl 4" Dorothea Haueser). (5) In a series of specimens collected 

 by Betty Knight near Paso Robles, pubescent pedicels are associated with fertile flowers and 

 glabrous pedicels with sterile flowers. This statement is more or less true of most material of this 

 variety. 



The upper pair of stamens are united by their filaments to the summit or they are free for 

 nearly a line's length ; in the lower pair the filaments are completely united or united for % or % 

 their length. There is in this matter of degree of union of the filaments obvious variation as in 

 S. coulteri. While the stigmas are conspicuously long in all plants of this form, this character is 

 one of degree only and will not serve ta this case for specific distinction. The Paso Robles plants 

 are, therefore, reduced to varietal rank. In aspect they are quite like S. coulteri. 



Refs. — Streptanthus coulteri Greene, Fl. Fr. 257 (1891) ; Jepson, Man. 416, fig. 408 

 (1925). Caulanthus coulteri Wats. Bot. King 27 (1871), type from Cal., Coulter. Our present 

 knowledge of the distribution of this species indicates that the Santa Clara Valley at or near 

 Saugus is the only station crossed by the route of Thos. Coulter, Var. lemmonh Jepson. Strep- 

 tanthus lemmonii Jepson, Man. 416 (1925). Caulanthus lemmonii Wats. Proc. Am. Acad. 23:261 

 (1888), type loc. "Cholame," Lemmon. S. parryi Greene, Fl. Fr. 257 (1891), type a spm. col- 

 lected by Parry, probably near Paso Robles. 



11. S. inflatus Greene. Squaw Cabbage. Desert Candle. Annual; stem 

 simple, very stout, becoming strongly hollow-inflated, 1 to 2i^ feet liigh; herbage 

 glabrous (rarely a little hispid at base) ; blades of cauline leaves erect, oblong- 

 ovate to elliptic or oblong, obtuse or merely subacute, entire or denticulate, sessile- 

 auriculate or -cordate, li^ to 10 inches long; racemes at first very dense; pedicels 

 glabrous or hispid, 2 to 4 lines long; flowers ascending, spreading or almost hori- 



