MUSTARD FAMILY 29 



Sandy bottoms or rocky places, 3000 to 4000 feet : upper Sacramento River to 

 Trinity Co. June. 



Locs. — Sisson, Cong don; Hayfork Mt., Tracy, 6453. 



Eefs. — Streptanthus barbatus "Wats. Proc. Am. Acad. 25:125 (1890), type loc. upper Sac- 

 ramento Eiver, Wilkes Exped.; Gray, Syn. Fl. 1^:169 (1895) ; Jepson, Man. 417 (1925). S. tor- 

 tuosus Gray; Torr. Phanerogamia Pac. Coast N. Am. 227 (1874), not Kellogg (1863). Cartiera 

 barbata Greene, Lflts. 1:226 (1906), 



14. S. cordatus Nutt. Stems one or few from a perennial root-crown, often 

 stout, 1 to 3 feet high; blades of basal leaves broadly spatulate-obovate, denticulate 

 or dentate, especially toward the apex, and often setose-ciliate, y2 to II/2 inches 

 long, on petioles i/4 to as long; blades of cauline leaves cordate-ovate to -oblong, 

 mostly obtuse and entire, sessile, 1 to 2i/4 inches long; flowers 6 lines long; sepals 

 at first greenish, turning purplish, of one size, erect, bearing a tuft of 3 or 4 short 

 bristles near the apex; petals purple, narrowly white-margined, li/^ to l^^ times 

 as long as the sepals, with ovate claw contracted above to a ligulate limb; pods 

 ascending or spreading, 2 to 3^4 inches long, IV2 to 2^/2 lines broad; pedicels short, 

 2 to 4 lines long; stigma subsessile; seeds orbicular, winged. 



Dry sandy loam or rocky slopes of mountains and montane valleys, 5000 to 

 7500 feet : Alpine Co. to Shasta and Modoc Cos. East to Colorado, north to Oregon. 

 June-July. 



Tax. note. — E. L. Greene (Pitt. 3:227) argues that the "eastern California" plants long 

 referred to Streptanthus cordatus do not represent Streptanthus cordatus Nutt., the original of 

 which was collected by Nuttall in the northern Rocky Mts. In the absence of a type specimen he 

 receives Wyoming and eastern Utah plants as true S. cordatus and points out that in these the 

 sepals are not bristly at tip as in the plants of eastern California. However, in a sheet of Strep- 

 tanthus cordatus, a Jones collection from Deep Creek, Utah, which is surely conspecific with our 

 plants of northeastern California, the calyces of one individual show this feature of a bristle- 

 tipped calyx, while in the other individual the sepals are glabrous. 



The brief original description of Streptanthus cordatus Nutt. applies to our California plants 

 save in that the flowers are described as greenish-white and the pods as deflexed. This statement 

 as to color may for various reasons be unimportant, but the second point is more difficult. While 

 the pods in our California plants are ascending or spreading, sometimes they are recurved as in 

 the somewhat immature specimens of M. S. Baker from the Tamarack road, eastern Shasta 

 County. It may be noted by way of comparison with other species that while the pods are typi- 

 cally reflexed in S. coulteri Greene, they are sometimes erect. Nuttall's original description cov- 

 ers explicitly the marked differences as to shape, petiolation and margin between the basal leaves 

 and the cauline ones. We know of no other Streptanthus in which just this kind of heterophyllism 

 occurs. 



Geographic considerations, too, lend some weight to the view that the plants on the western 

 edge of the Great Basin may be the same as Nuttall's original plant from the north-easterly side 

 of the Great Basin. If the type specimens are not extant, as said, the doubts raised by Greene 

 may never be resolved, though a careful survey of the Streptanthi along the path traversed by 

 Nuttall in crossing the northern Rocky Mts. might furnish a solution. In this work we retain 

 the well-known name for our plants as conspecific with the plants on the eastern side of the Great 

 Basin commonly referred to this species. 



Streptanthus cordatus in California occurs from Alpine Co. to Modoc Co. This form is well- 

 known. S. cordatus is little known to us from "eastern California southward" (Pitt. 3:227) 

 save in modified and very rare states which we specially indicate below as varieties. Citation of 

 characteristic specimens of the species follows here. 



Locs. — Ebbets Pass, Brewer; Red Clover Creek, Plumas Co. ; summit Tamarack road between 

 Clover Creek and Burney Valley, M. S. Baker; Davis Creek, Modoc Co., Austin ^ Bruce 2245; 

 Ft. Bidwell, Manning 389 ; Mt. Bidwell, Jepson 7896 ; Pine Creek, Warner Range, L. S. Smith 965. 



Two quite unlike high altitudinal California forms of this species occur southward as f oUows : 

 Var. exiguus Jepson var. n. Dwarfish, 4 to 7 inches high; leaves % to l^/^ inches long, narrowly 

 lanceolate, entire, acute or acutish, the cauline merely auriculate-sessUe ; flowers 4 to 5 liaes long. 

 — (Plantae reductae, unc. 4-7 altae; folia unc. %-l% longa, anguste lanceolata, integra, acuta 

 vel acutiuscula, caulina auriculato-sessilia; flores lin. 4-5 longi.) — Sonora Pass, 10,000 feet. 

 Brewer 1885. Var. duranii Jepson var. n. Stems 14 inches high ; leaves as in the species ; racenie 

 loose ; flowers 4 to 5 lines long ; sepals deep purple, a little loose or tending not to overlap by their 

 edges (whereas in the species there is a regular or even overlapping of the sepals by their edges) ; 



