36 



CRUCIFERAE 



1. T. integrifolium Endl. Stem simple or branched only above, 3 to 6 feet 

 hiu'h; liorbajxo ^Wahrous; blades of basal leaves ovate-lanceolate, entire or shallowly 

 n'pai'ul-dentate, 1 to 6 inches lonp:, drawn down into petioles nearly as lonp:; blades 

 of npper leaves linear-lanceolate, sessile (the lower ones sometimes auricnlate), 

 acnte, erect; flowers crowded in subcapitate racemes; racemes 1 to 2 (or 4) inches 

 Ion?, several in a terminal sli«rhtly leafy-bracteate panicle; pedicels 1 to 3 lines 

 Ion J'; racemes little elonjjated in fruit, the pods therefore crowded and forming a 

 sort of "brush"; pods widely spreading, slender, torulose, curved upwards, 1 to 

 iVs inches long; stipe i/4 to V^ line long. 



Moist situations or in stream bottoms, 2300 to 8000 feet : Mohave Desert; White 

 Mts. East to Colorado, north to eastern Washington. June-Aug. 



Note on the glands. — Tn Thelypodium integrifolium, as it occurs in Oregon and Washington 

 (the tvpe region), the flowers have four distinct nectar glands. In Mohave Desert plants (rep- 

 resenting the form called T. affine Greene) the flowers have the 4 glands united in such various 



degrees that they may occur as two only and entire, 

 or two and variously notched as shown in fig. 131. 

 i )\ /^ )\\ A I \\ fl i\ ^^^ further, under "Ix)cs." below, Parish 1485. This 



AJi) JAjUH. ill a lit II i 11 condition of more or less partial union of the glands 



is to be seen also in plants at Wadsworth, Nev. 

 (Palmer 22). Thelypodium affine Greene, consid- 

 ered as distinct from T. integrifolium, rests essen- 

 tially upon the gland character, but we regard this 

 character as variable and uncertain. In aspect and 

 in detail of structure the Californian plants seem 

 essentially like those of the north, that is, those of 

 Oregon and Washington. 



Locs. — So far as now known to us, this species 

 occurs in California only in the White Mts. (Eob- 

 erts Ranch, Wyman Creek, V. Duran 1688) and in 

 a crescent-shaped area on the westerly margin of the 

 Mohave Desert along the base of the mountains or 

 on their lower slopes as follows: Tehachapi Mts.; 

 Lancaster, Parish ms; Rancho Verde, VictorvUle, 

 Parish 10,532; Rabbit Sprs., S. B. <S' W. F. Parish 

 1485 (glands mostly 2 and entire, in some cases notched, in at least one flower 4 distinct glands) ; 

 Cushenbury Sprs., Parish ms. Palmer (Contrib. U. S. Nat. Herb. 1:7) also found it at Victorville 

 and says the leaves were gathered by the Indians and eaten after being cooked. 



Refs. — Thelypodium integrifoliitm Endl.; Walp. Rep. 1:172 (1842); Jepson, Man. 412 

 (1925). Pachypodium integrifolium Nutt. ; T. & G. Fl. 1:96 (1838), type loc. Rocky Mt. plateau 

 towards the Columbia River as far as Walla Walla, Nuttall; H. & A. Bot. Beech. 321, t. 74 (1840). 

 Pleurophragma integrifolium Rydb. Bull. Torr. Club 34 :433 (1907) . T. affine Greene, Pitt. 4 :314 

 (1901), type loc. Tehachapi, Greene. 



2. T. laciniatum Endl. Stem stout, freely branching, 1% to 2l^ feet high; 

 herbage glabrous; leaf -blades deltoid-ovate to oblong or oblong-lanceolate, irregu- 

 larly segmented and toothed, a little fleshy, ll^ to 8 inches long, the petioles i/^ to 

 as long; racemes very dense, 1/2 to IVs feet long; flowers 4 to 5 lines long; petals 

 white, linear or oblong, much exserted; stamens exserted; pods very slender, spread- 

 ing or horizontally divaricate, VA to 6 inches long; pedicels 1 to 2 lines long. 



Valleys and foothills, 2000 to 4000 feet : Inyo Co. ; Siskiyou Co. North to west- 

 em Nevada and eastern Oregon and Washington. May-June. 



Tax. and geog. note. — The very long and dense fruiting racemes of slender elongated pods 

 is a feature of this striking but somewhat polymorphous species. A detail relates to the fruiting 

 style which we find somewhat globose-thickened under the stigma. The species occurs in Cali- 

 fornia sparingly, and we have records only as follows: Big Pine, Eeller 8262; Yreka (Ann. Mo. 

 Bot. Gard. 9:273). 



Var. milleflorum Payson. Leaves tending to be subentire or merely dentate ; racemes with 

 ascending pedicels, flowers and pods. — Northern part of the Great Basin, entering California in 

 eastern Sierra Co. (Sierra Valley, ace. Payson, Ann. Mo. Bot. Gard. 9:275). 



Refs. — THELYPODrUM LACINIATUM Endl.; Walp. Rep. 1:172 (1842); Jepson, Man. 412 

 (1925). Macropodium laciniatum Hook. Fl. Bor. Am. 1:43 (1829), type loc. Wallawalla and 



Fig. 131. Thelypodium integrlfolium 

 Endl. Variation in the lobation of the 

 flower glands: a, 6, c, Victorville, Parish 

 10,532; d, e, Victorville, Johnston \112\ 

 f, g, Wadsworth, Nev., Palmer 22. 



