38 CRUCIFERAE 



or shallowly (or sometimes sinuately) pinnatifid witli a few coarse salient lobes at 

 the middle and below, 2 to 7 inches lonj?, contracted to a petiole Yo to 2 inches long; 

 blades 111' upper leaves lance-linear, attenuate at both ends, sessile; racemes in 

 anthesis somewhat dense, soon becoming loose; flowers 2 to 2Vi> lines long, the parts 

 spreading, the sepals and petals nearly or quite rotate; sepals dark lavender with 

 white margins, carinate, hooded at apex; petals pale lavender with deeper veins, 

 obovate or cuneate-oblong, only slightly longer or slightly shorter than the sepals; 

 anthei-s yellow or purple; capsule slender or subfiliform, 2 to 214 inches long; 

 pedicels glabrous; stigma small, 2-lobed. 



Open hill slopes or plains in heavy black adobe or friable gray clays, 700 to 3000 

 feet : inner South Coast Range from western San Joaquin Co. to southeastern San 

 Luis Obispo Co. Mar.-June. 



Note on relationship. — Thelypodium lemnionii, an excellent species, has been somewhat rarely 

 collected. It grows in colonies of about one-quarter to three acres usually, which favor the friable 

 clays of steep hill slopes. The colonies are commonly rather densely populated and rather well 

 defined. It seems likely that they are on the whole well isolated with reference to each other. 

 We have observed them only at intervals of about six to twenty miles. 



While differing very strongly from Thelypodium laciniatum, the type of the genus, Thely- 

 podium lemmonii seems nevertheless more at home in Thelypodium than in Streptanthus. The 

 petals are nearly rotate, while the linear sepals are exactly and somewhat stiffly rotate, filling the 

 intervals between the petals. The flower is thus not at all streptanthoid, either considered as a 

 whole or in relation to the petals which are nearly plane or only slightly cupped, not at all crisped 

 and without any differentiation into claw and limb. On tlie whole the flowers are fairly thely- 

 podioid, especially as compared with tliose of Thelypodium brachycarpum Torr. The filaments 

 are terete, quite distinct and slightly fleshy; they are rarely a little hairy. In one colony, as 

 observed in the Yeguas Hills, where the plants are all of one aspect and habit, with uniform leaf, 

 raceme and flower characteristics, some flowering axes show closely erect pods, other axes show 

 reflexed pods, other axes still show spreading pods (Jepson 16,203). The flowers are sometimes 

 obviously a little "physiologically bilabiate" as to position of the petals which stand in pairs: 

 those of the lower pair spread a little right and left, those of the upper pair are slightly divergent 

 and turned backward. A colony of these plants is slightly fragrant during anthesis. 



Locs. — Tracy (Fl. Fr. 263) ; Zapato Chino Creek, sw. Fresno Co., T. Brandegee; Diablo 

 Range, sw. Fresno Co., Jepson 15,391; Cholame, Lemmon 4572; Estrella plain. Barber; upper 

 Waltham Creek, w. Fresno Co., Jepson 16,160 ; Palo Prieto Canon, e. San Luis Obispo Co., Jepson, 

 16,198; Yeguas Hills, n. of Carrizo Plain, Jepson 16,203; Sumner ranch, Bitterwater Creek, 

 Temblor Eange, Jepson; Carrizo Plain, opp. Elkhorn Scarp, Jepson 16,219; Cuyama, se. San 

 Luis Obispo Co., Eastwood. 



Refs. — Thelypodium lemmonh Greene, West Am. Sci. 3:156 (1887), type loc. Lemmon's 

 ranch, Cholame, San Luis Obispo Co., Lemmon; Fl. Fr. 263 (1891) ; Gray, Syn. Fl. 1:178 (1895) ; 

 Jepson, Man. 412 (1925). Caulanthus anceps Payson, Ann. Mo. Bot. Gard. 9:303 (1922). 



5. T. lasiophyllum Greene. Stem simple, or branching above, y2 to 6 feet 

 high; herbage hispid with scattered hairs or nearly glabrous above; blades of lower 

 leaves sinuately pinnatifid with mostly acute denticulate or entire segments, 2 to 

 10 inches long, the upper lanceolate, less lobed or merely denticulate, all petioled, 

 or the upper rarely sessile; raceme very densely flowered, much elongated in fruit; 

 pedicels 1 line long; petals white, yellowish, or sometimes roseate, 1% to 2 lines 

 long; sepals oblong, scarcely the length of the narrow petals; pods straight or some- 

 what curved, strictly defiexed, 2 to 4 inches long, ^ to ^ line wide. 



Open foothills, good soil, 50 to 2100 feet : coastal Southern California; Colorado 

 and Mohave deserts; Coast Ranges and their bordering plains, mostly toward the 

 interior. North to Washington. Apr. 



Variation and geographic range. — This species is fluctuatingly variable in nearly every gross 

 character. The plants vary greatly in size and are especially luxuriant on "burns." The leaves 

 are notoriously variable and exhibit a wide range in pinnate division; they are glabrous to hirsute 

 and vary greatly in size. The flowers vary in color. The petals show a range of variation from 

 oblong-spatulate to linear in plants growing side by side (Howell Mt., Tracy 1500, 1500^^). The 



