MUSTARD FAMILY 39 



pods vary from very slender to stout, from straight to much curved, assume very diverse positions, 

 and vary much in length. These variations, hov^ever, occur without any apparent or definite 

 relation to the geographic distribution of the species in the California coastal region. 



While most Thelypodium species are inclined to be infrequent in individuals, this species is 

 one of the most common and widely distributed native plants of its immediate alliance in the 

 Coast Range hills, especially toward the inner range, but has never been collected in the Sierra 

 Nevada foothills north of Tulare Co. so far as known to us. It extends south to San Diego Co. 

 and in a slightly or obscurely modified form through the Mohave Desert eastward to Utah. Like- 

 wise it ranges north to Washington. Its preference for the Coast Ranges as against the Sierra 

 Nevada foothills is all the more remarkable in that it extends so far beyond the Coast Ranges 

 proper to the north and to the south without obvious modification. It may be hazarded that 

 slightly higher winter or spring temperatures in the Sierra Nevada foothills account for its 

 absence in that range. The varieties named below are of little importance morphologically. 



Locs. — S. Cal. : Vallecito, Jepson 8598 ; Blair Valley, e. San Diego Co., Jepson 8676 ; Cuya- 

 maca Mts., Newlon 359; Campo, Abrams 3573; San Diego, Jones 2634; Painted Canon ne. of 

 Mecca, Jepson 11,657; Arroyo Seco, Los Angeles, Peirson 327; Hesperia, Newlon 469; Antelope 

 Valley, Davidson; Mohave sta.. Heller 7751; Ord Mt., Jepson 15,504. Coast Ranges: Zapato 

 Chino Creek, sw. Fresno Co., Jepson 15,365; Paso Robles, Barber; Carmel, Heller 6586; Crystal 

 Sprs., San Mateo Co., C. F. Baker 459 ; Los Buellis Hills, E. J. Smith ; Berkeley, Tracy 599 ; Liver- 

 more, Bioletti; Mt. Diablo, Jepson 9219; Briones Hills, Chandler; Montezuma Hills, Solano Co., 

 Jepson; Gates Canon, Vaca Mts., Jepson 13,349; Napa Range e. of Calistoga, Jepson 13,347; 

 Santa Rosa, M. S. Baker; South Mill Creek, e. of Ukiah, Jepson 9225; Blue Lakes, Lake Co., 

 Jepson 13,350. West side of the Great Valley: Maricopa, Kern Co., Jepson 12,166; Huron, w. 

 Fresno Co., C. F. Meyer 266 ; Bethany, w. San Joaquin Co., C. F. Baker 2791 ; Colusa, Chandler 807. 



Var. inalienum Rob. Pods slender, erect or spreading. — Solano Co, to San Luis Obispo Co. 

 as follows: VacaviUe, Jepson 13,352, 13,353; Livermore, Greene; San Luis Mt., Summers. This 

 variety differs from var. rigidum because of its slender pods and from the species because of its 

 erect pods, but these characters, looking toward the species, are inconstant. 



Var. rigidum Rob. Plant rigid, often glabrous ; pods stout, rigid, deflexed-spreading, some- 

 what spinescent; pedicels V^ line long. — Glenn Co. to eastern Contra Costa Co. as follows: Wil- 

 lows, Jepson, 13,351; Main Prairie, Solano Co., Jepson 13,345; Livermore, Greene. Pods often 

 broadest at base and tapering to apex; they may be ascending, spreading and descending on 

 one raceme. 



Var. utahense Jepson. Leaf -blades thin, rather more glabrous than in the species, the lobes 

 rounded (mostly acute in the species); fruiting raceme sparse; pods slender, very rigidly and 

 angularly divaricate, and straight or sometimes a little curved. — -Colorado and Mohave deserts. 

 East to Utah. This is an inconstant form which is scarcely tenable. The specimens representing 

 it vary amongst themselves about as much as they do collectively from the species. Intermediates 

 to the species occur in the Colorado Desert. It is not uncommon in the deserts. Eastern San 

 Diego Co.: Vallecito, Jepson 8574, 8599; Mason Valley, Jepson 8712; Blair Valley, Jepson 8677c; 

 Wagon Wash near Sentenac Canon Jepson 12,477. Utah: St. Thomas, Goodding 700. 



Refs. — Thelypodium lasiophyllum Greene, Bull. Torr. Club 13:142 (1886), Man. Reg. 

 S. F. Bay 19 (1894) ; Jepson, Fl. W. Mid. Cal. 212 (1901), ed. 2, 180 (1911), Man. 412, fig. 406 

 (1925). Turritis lasiophylla H. & A. Bot. Beech. 321 (1840), type from Cal., Douglas. Guillenia 

 lasiophylla Greene, Lflts. 1 :227 (1906). Sisymbrium reflexum Nutt. Jour. Acad. Phila. n.s. 1 :183 

 (1847), type loc. near San Pedro, Cal., Nuttall, and Proc. Phil. Acad. 4:25 (1848) ; B. & W. Bot. 

 Cal. 1:41 (1876). Var. inalienum Rob.; Gray, Syn. Fl. r:177 (1895), type loc. Nipoma, 

 Brewer 417; Jepson, Fl. W. Mid. Cal. 212 (1901), Man. 413 (1925). Guillenia inaliena Greene, 

 Lflts. 1:228 (1906). Sisymbrium acutangulum B. & W. Bot. Cal. 1:41 (1876), not DC. Var. 

 RIGIDUM Rob.; Gray, Syn. Fl. 1»:177 (1895) ; Jepson, Fl. W. Mid. Cal. 212 (1901), Man. 413 

 (1925). T. rigidum Greene, Pitt. 1:62 (1887), type loc. Antioch, Greene. Guillenia rigida 

 Greene, Lflts. 1 :228 (1906). Var. utahense Jepson, Man. 413 (1925). T. utahense Rydb. Bull. 

 Torr. Club 29:233 (1902), type loc. St. George, Utah, Jones in 1880. 



6. T. longirostris Jepson. Stem much-branched, 1 to 1% feet high; herbage 

 glabrous and glaucous; leaf -blades lanceolate to linear-lanceolate, the lower repand- 

 dentate, the upper entire, all contracted to a short petiole, the whole 1 to 2^2 inches 

 long; flowers 2 to 2% lines long, pale yellow or pinkish, soon reflexed, borne on slen- 

 der pedicels 1/2 to 1 line long; sepals green, or somewhat purplish-tinged, scarious- 

 margined ; petals pale yellow or pinkish, narrow, limb very short, narrower than 

 the expanded summit of the claw; pods curved or straight, divaricate or deflexed, 

 nerved, 1 to l^/^ inches long, 14 to % line wide, tapering to a beak 1 to 3 lines long; 

 seeds winged. 



