58 CRUCIFERAE 



Dcntaria intcgrifolia (including its varieties and more especially the var. californica) is 

 extroiiiely varialilo in foliage and in habit. I^eaf or leaflet outline is especially fluctuating. 

 A single individual often exliibits five or six distinct shapes of leaflets, while it is not infrequently 

 one's experience that fifteen or twenty more or less marked leaf variants can be had from a series 

 of plants froiM oiu' station, the plants otherwise not dissimilar. 



Var. californica Jepson. Leaves or leaflets larger and thinner, as a rule, than in the species; 

 corolla white or pale rose-color. — Shady banks or in rich woods or tliickets, 50 to 8700 feet, 

 throughout eismontanc C:Uifornia. 



Geog. distribution. — This form, so ncarlj' related to the species, grows in the Avooded canons 

 of hills bordering the open wet valley flats indicated above as occupied by D. intcgrifolia but is 

 more widely distributed by far and is common everj^herc in the hill country beyond the range 

 of D. intcgrifolia. Its leaves were described by Nuttall as crenate or incisely denticulate, and 

 are in any event more commonly denticulate than in D. intcgrifolia, and are sometimes pale purple 

 beneath. In the main its characters suggest tliat it is an edaphic variant. A long series of speci- 

 mens of var. c-alifornica may be cited as follows. S. Cal. : San Diego, G. W. Dunn; Arroyo Scco, 

 Los Angeles, Brmmton 789; Pacoima Canon, San Gabriel Mts., Peirson 265; Lady's Harbor, 

 Santa Cruz Isl., Mabel Peirson. Coast Ranges: San Luis Obispo, Ida Blochman; Los Altos (hills 

 back of), Santa Clara Co., Jepson 9103; Arroyo Mocho, Jepson 10,677; Mt. Diablo, Jepson 9850; 

 Berkeley Hills, Earl Mulliken 4; Koss Valley, Jepson 30k; Papermill Creek, Marin Co., Jepson 

 10,311a'; St. Helena, Jepson 32e; Knights Valley, Jepson 32d ; Duncan Mills, Davy 1644; Na- 

 varro, Mendocino Co., Byxbee; Kneeland Prairie, Tracy 4878; Trinity Summit, Davy 5832; 

 Marble Mt., Jepson 2844; Crescent City, Howell. Sierra Nevada: Tule River, Tulare Co., Purpus 

 5075; Yankee HUl, Tuolumne Co., A. L. Grant 650; New York Falls, Amador Co., Hansen 578; 

 Big Mdws., Plumas Co., B. M. Austin; Little Chico Creek, E. M. Austin 332; Cow Creek Canon, 

 Shasta Co., M. S. Baker; Shasta Sprs., Butler 649. 



Var. tracyi Jepson. Leaves a very dense deep purple beneath, the basal smaller than in var. 

 californica; flowers deep rose-pink, slightly smaller. — Woods, Van Duzen River, Humboldt Co. 



Var. pachystigma Wats. Raceme sessile or nearly so. — Plumas Co. 



Refs.— Dentabia integrifolia Nutt; T. & G. PL 1:88 (1838), type loc. Monterey, Nuttall; 

 Jepson, Fl. W. Mid. Cal. 221 (1901), ed. 2, 188 (1911) ; Man. 426 (1925). Cardamine integri- 

 folia Greene, Bull. Cal. Acad. 2:389 (1887). Var. californica Jepson, Fl. W. Mid. Cal. 222 

 (1901), ed. 2, 188 (1911), Man. 426 (1925). D. californica Nutt. I.e., type loc. Monterey, Nut- 

 tall; Wats. Proe. Am. Acad. 14:289 (1879); Bot. Cal. 2:430 (1880); Gray, Syn. Fl. r:159 

 (1895). Cardamine californica Greene, Fl. Fr. 266 (1891). C. puucisecta Benth. PI. Hartw. 297 

 (1848), type loc. Monterey, Hartweg 174; B. & W. Bot. Cal. 1:30 (1876). C. cuneata Greene, 

 Bull. Cal. Acad. 1:74 (1885), tvpe loc. Jolon, Monterev Co., Greene. D. cuneata Greene, Pitt. 

 3:123 (1896). Cardamine nuttaUii Greene, Bull. Cal. Acad. 2:389 (1887), Fl. Fr. 266 (1891). 

 C. sinuata Greene, Erythea, 1:148 (1893), type loc. Crescent City, Thomas Howell. D. sinuata 

 Greene, Pitt. 3:123 (1896). Var. tracyi Jepson, Man. 426 (1925), type loc. Blue Slide, Van 

 Duzen River, Tracy 5426. Var. pachystigma Wats. Proc. Am. Acad. 14:289 (1879), type loc. 

 Plumas Co., Lemmon, Fl. M. Austin, Ames; Jepson, Man. 426 (1925). D. pachystigma Wats.; 

 Gray, Syn. Fl. P:155 (1895). 



4. D. macrocarpa Nutt. Oregon Toothwort. Stem 4 to 8 inches high, aris- 

 ing from elongated thick rootstocks; leaves pinnately divided into 5 linear or oblong 

 lobes % to 1 inch long, or sometimes palmately parted; cauline leaves 1 to 3; raceme 

 nearly sessile; petals pink; pods 1 to IV2 inches long; styles in fruit 4 to 5 lines long. 



Shady slopes, 10 to 4000 feet : Plumas Co. to Del Norte Co. North to British 

 Columbia. Mar.-Apr. 



Locs.— Plumas Co. (Sjti. Fl. l':154); Humbug Mt., Siskiyou Co., Butler 1152; Crescent 

 City, Howell 109. 



Refs. — Dentaria macrocarpa Nutt.; T. & G. Fl. 1:88 (1838), type loc. Columbia River, 

 Nuttall; Jepson, Man. 427 (1925). C. pulcherrima Greene, Erythea 1:148 (1893), type loc. 

 Mosier, e. Ore., Howell. 



15. SMELOWSKIA C. A. Mey. 



Low caespitose perennials with stellate-pubescent herbage. Leaves deeply pin- 

 natifid or bipinnatifid. Flowers in racemes, small, white, yellowish or purplish- 

 tinged. Anthers slightly sagittate at base. Pod oblong-lanceolate or lanceolate, 

 more or less obcompressed, the valves strongly keeled; style short. — Species 7, 

 western North America and central Asia. (Timotheus Smielowski, Russian botan- 

 ist, 19th century.) 



