118 SAXIFRAGACEAE 



SaxifrnKft (Bull. Torr. Club 23:366, — 1896). Greene, E. L., Two Californian Saxifrages (Bull. 

 Torr. Club 23:2.') — 1896). Gunthart, A., Bcitriigo zur Bliitonbiolopic dcr Gattunp Saxifraga 

 (Bibiio. Bot. ll"':63-94, — 1902). Enfjlcr, A., and Irinschcr, E., Saxifragacoae-Saxifraga (Eng- 

 ler, Pflzr. 4'": 1-709, figs. 1-124,-1916). Engler, A., Saxifraga (Engler, Pflzr. 4'": para gen. 

 1-47, figs. 1-4, — 1918). Johnson, A. M., Hevision of the N. Am. species of the sect. Boraphila, 

 genus S.'ixifraga (Univ. Minn. Stud. Biol. Sci. 4:1-109, pis. 1-19,-1923) ; Studies Ln Saxifraga. 

 I. (Am. Jour. Bot 14:323-326, pis. 40-47,-1923) ; II. (I.e. 18:797-802, pis. 53-54,-1931). 



Stems with the leaves all basal or nearly so. 



Inflorescence without bulblcts; perennials. 

 Leaves not orbicular or cordate. 

 Petals not spotted. 



Leaves commonly glabrous, at least on upper surface; petioles mostly short. 

 Calyx-lobes erect. 



Plants 1 to 3 feet high ; flowers in an elongated panicle..!. S. oregana. 

 Plants 2 to 14 inches high. 



Scapes glabrous except at base; flowers in a single terminal 



capitate cyme 2. S. aprica. 



Scapes glandular-pubescent; cymules in a commonly narrow 



panicle, rarely capitate 3. S. nidifica. 



Calyx-lobes reflexed; cymules in a loose panicle i.S.fall<ix, 



Leaves commonly with soft scattered hairs on the upper surface; petioles ^/4 

 to as long as or longer than the blades; flowers in an open panicle; 



calyx-lobes reflexed or becoming reflexed 5. S. calif ornica. 



Petals with 2 yellow-green spots near the base ; filaments petaloid....6. S. marshallii. 

 Leaves orbicular or cordate; petiole longer than the blade. 



Stems not scapose, the stems commonly 1 or 2-flowered 7. S. rivularis. 



Stems scapose, bearing many-flowered panicles. 



Scapes from a more or less horizontal rootstock; leaves simply crenate or 



serrate 8. S. puTictata. 



Scapes from an erect bulb-like scaly root-crown ; leaves doubly serrate 



9. S. mertensiana. 

 Inflorescence with bulblets; high montane. 



Perennial; stoutish plant 6 to 11 inches high; rare in Cal 10. S. hongardii. 



Annual; delicate or slender plant 2 to 8 inches high, frequent 11. S. bryophora. 



Stems densely and equably clothed with linear leaves; caespitose alpine plants 12. S. tolmiei. 



1. S. oregana Howell. Marsh Saxifrage. Scapes stoutish, 12 to 22 inches 

 higli ; stems pubescent, the upper part glandular ; leaf-blades obovate to oblanceo- 

 late-spatulate, 2 to 7 inches long, narrowed at base to a margined petiole, mostly gla- 

 brous, sometimes ciliate; flowers in small thick clusters in a narrow thyrse-like 

 panicle; petals obovate to oblong-obovate, 1% lines long. 



Swamps or mar.shy meadows, 3500 to 7400 feet : Sierra Nevada from Tulare 

 Co. to Siskiyou Co. North to Washington. June- Aug. 



Tax. note. — These plants of the Sierra Nevada montane meadows, hitherto called Saxifraga 

 integrifolia Hook. (Fl. Bor. Am. 1:249 t. 86, — 1834), a low valley species of Oregon and Wash- 

 ington, are here referred to S. oregana Howell. In S. oregana the leaves are larger, widest to- 

 wards the apex and taper insensibly or abruptly to the base, whereas in S. integrifolia the leaves 

 are broadest at base and tend to be ovate or oblong, so that the base of the blade is usually 

 rounded or obtuse. The rootstock of S. oregana is, furthermore, heavier and the leaf-bases are 

 broad and thin and do not develop in age a fine somewhat wire-like thatch at base of stem as in 

 S. integrifolia. While the leaves in S. oregana vary from entire or denticulate to markedly 

 serrate, and from forms with a long petiole-like base to subsessile or sessile, such range of varia- 

 tion is, however, not significant and represents a verj' ordinary sort of variation in many Saxi- 

 fraga species. The var. sierrae (Gov.) with "sessile" leaves we hold an inconsiderable form. 



Logs. — Freeman Creek, Tulare Co., Jepson 4885; Giant Forest, K. Brandegee ; Horse Corral 

 Mdw., King's River, Jepson 768; Huckleberry Creek, Huntington Lake, Jepson 13,076; Straw- 

 berry, South Fork Stanislaus River, Jepson 6524; Kennedy Lake, Tuolumne Co., A. L. Grant 

 330; Myers sta., Eldorado Co., Ottley 960; Colby, Butte Co., E. M. Austin; ridge n. of Hot 

 Springs Valley, Lassen Peak, Jepson 12,297; Pine Creek, Lassen Co., M. S. Balcer; Bear Valley 

 Mts., Modoc Co., If. S. Balcer 564; Twelve-mile Creek, e. side Mt. Bidwell, Manning; Deadhorse 

 Canon, Siskiyou Co., M. S. Balcer; Sisson, Jepson 13,476 ; Marble Mt., Siskiyou Co., Chandler 1548. 



Refs. — Saxifraga oregana Howell, Erythea 3: 34 (1895), "mountain marshes of Oregon", 

 Howell. S. integrifolia Jepson, Man. 455, fig. 446 (1925), not Hook. Micranthes integrifolia 

 Small, N. Ajn. Fl. 22:137 (1905) in part. S. integrifolia var. sierrae Gov. Proc. Biol. Soe. Wash. 



