SAXIFRAGE FAMILY 157 



calyx greenish, dull white, or sometimes purplish-tinged, glabrous, sometimes mi- 

 nutely puberulent, sometimes the lobes tipped with a few pilose hairs; petals 

 white, 1/4 to 1/^ as long as the stamens ; ovary glabrous, or sometimes puberulent, 

 but covered with rather long spines, these rarely interspersed with shorter gland- 

 tipped ones ; berry usually small, 3 to 4 or 5 lines in diameter, the spines slender. 

 Hills or narrow canon flats, most characteristic of dry exposed or rocky slopes, 

 300 to 2500 feet: central Coast Ranges from Monterey Co. to Mendocino Co. 

 Jan.-Apr. 



Variability. — Of all our Californian species of gooseberries, none perhaps is more variable 

 than R. californieum H. & A. except E. menziesii Pursh. In its region every canon, every flat, 

 every hillslope with a definite topographic character has its own type of shrub. The plants differ 

 in spininess, in the extent to which their branchlets are zig-zag, in degree of pubescence of leaves, 

 of peduncles, of calyces, of ovaries, and in number of spines on the fruit. The flowers are seldom 

 absolutely glabrous, frequently almost so, though always inconstant in a minute manner; while 

 thus markedly varying in degree, the pubescence is usually extremely scanty, though exceptionally 

 one finds rather pubescent flowers. The fruits show the greatest range in the matter of presence 

 of spines. Shrubs at Lake San Andreas (Jepson 9540) show quite unarmed berries as do shrubs 

 at Long Eidge road near "Wrights, Santa Cruz Mts. (Wolley-Dod 402). From this quite naked 

 condition we have shrubs exhibiting just a few scattered spines on the berry (R. oligacanthum 

 Eastw.) continuously to those which develop rather densely spiny fruits. 



The original specimens of E. californieum H. & A. (California, 1831) were gathered by 

 David Douglas in early flower with few leaves as yet developed, and represent the thorny rigid- 

 stemmed quite glabrous form of rocky hills having glabrous non-glandular foliage and flowers 

 and moderately armed ovaries with short spines. The original Douglas specimens of E. occi- 

 dentale H. & A., collected in California in 1831, are in a more advanced stage; they are like the 

 E. californieum specimens in every technical particular, differing only in aspect, the branches 

 being leafier and more pliable. Our examination of the types of these forms was made at the 

 Eoyal Botanic Gardens in Kew, England, in 1930, some forty modern specimens from west cen- 

 tral California being critically compared with them. As to flower pubescence, our series of 

 specimens show every transition from branchlets bearing glabrous flowers and flowers with a few 

 hairs to specimens in which the flowers are obviously a little pubescent. 



Locs. — Priest Valley, San Carlos Eange, Jepson 2670; Guadalupe Mine, Santa Cruz Mts., 

 Jepson 9088 ; Los Gatos, Heller 7249 ; Saratoga, Pendleton 287, 519, 524, 527 ; Stanford, C. F. 

 Baker 316; Lake San Andreas, San Mateo Co., Jepson 9540, 9543; Pilarcitos Creek, San Mateo 

 Co., Davy 1058; Los Buellis Hills, Santa Clara Co., E. J. Smith; Arroyo Moeho, Mt. Hamilton 

 Eange, Jepson 10,682; Berkeley Hills, Jepson 6229b; Mt. Diablo; Seaview, Sonoma Co., M. S. 

 Baker; Watson Caiion, Sonoma Co., Bioletti; Calistoga (mts. e.), Jepson 13,511; Kelseyville, 

 Irwin; Potter Valley, Purpus 910, 1007. 



Eefs. — EiBES CALiFORNicuai H. & A. Bot. Beech. 346 (1840), type from Cal., Douglas (typ. 

 vidi) ; Jepson, El. W. Mid. Cal. 274 (1901), ed. 2, 202 (1911), Man. 473, fig. 469 (1925). Gros- 

 sularia calif arnica Gov. & Britt. N. Am. El. 22:214 (1908). E. occidentale H. & A. I.e., type from 

 Cal., Douglas (typ. vidi). E. oligacanthum Eastw. Proc. Cal. Acad. ser. 3, Bot. 2:246, pi. 24, 

 figs. 8a, 8b (1902), type loc. near Mansfield Eanch, King City, Eastwood (typ. vidi). 



21. R. victoris Greene. Victor's Gooseberry. Bush 2 to 5 feet high; nodal 

 spines 1 to 3, the newer branches prickly ; leaf -blades incisely 5-lobed, cordate at 

 base, crenate, the lower pair of lobes much smaller, sparsely puberulent, and with 

 gland-tipped hairs and sessile glands or dots, especially below; peduncles 5 to 6 

 lines long, 1 or 2-flowered; flowers 5 to 7 lines long; sepals greenish- white; ovary 

 thickly covered with short gland-tipped processes and a few longer non-glandular 

 spines. 



Canons, 500 feet: near Lake Lagunitas, north side of Mt. Tamalpais, Marin 

 Co., only one or two shrubs known. Mar. 



Var. greeneianum Jepson. Leaf -blades truncatish at base to cordate, not at all or scarcely 

 glandular, % to 1% inches wide, peduncles y2 to 1% inches long; flowers 6 to 8% lines long; 

 ovary densely covered with short-stiped glands (appearing sometimes as if tessellated), non- 

 glandular spines none; berry oval, yellow or red-brown, thickly covered with short gland-tipped 

 spines, 6 to 8 lines in diameter. — Canons or flats, 400 to 1500 feet: Hoods Peak Eange (Sonoma 

 Creek, Heller 5773, Adobe Canon, Michener (f- Bioletti; Niebaum's Dam, near St. Helena, Jepson 

 13,499) ; Napa Eange (Mt. St. Helena, Alice King, Middleton grade, Jepson 13,507) ; Vaca Mts. 

 (Gates Canon, Jepson 13,506, 13,508, 13,528). 



