166 ROSACEAE 



Rydb. Mem. N. Y. Bot. Gard. 1:206 (1900). P. acuviinatum Rydb. N. Am. Fl. 22:253 (1908), 

 type loc. Big Arroyo, Tulare Co., Culbertson 4540, herbage sparingly strigosc, calyx-lobes acumi- 

 nate, petals acute or acuminate. 



4. HOLODISCUS ^Faxim. 



Deciduous shrubs with toothed or lobed leaves and no stipules. Flowers creamy- 

 white, small, numerous in terminal panicles. Petals 5, rounded. Stamens 20, on 

 a ring-like perigynous disk. Pistils 5, distinct. Pods short-stipitate, hairy, 1- 

 seeded, tardily dehiscent or indehiscent. Ovules 2, pendulous. — Species 2, western 

 North America. (Greek holo, whole, and diskos, a disk, some related genera with 

 lobed disks.) 



1. H. discolor Maxim. Cream Bush. (Fig. 160.) Shrub 3 to 6 or sometimes 

 14 feet high; leaf-blades ovate to ovate-elliptic, coarsely serrate or incised above the 

 entire broadly cuneate (or sometimes subtruncatish) base, green and slightly 

 pubescent above, paler and pubescent beneath with soft hairs, % to 3 inches long; 

 petioles 2 to 6 lines long; panicles ample, 3 to 8 inches long, often half drooping in 

 anthesis; flowers 1^/^ lines long; follicles about 1 line long. 



Caiions in the hills, 5 to 5000 feet : along the coast from Los Angeles Co. to Siski- 

 you Co. ; occasional in the Sierra Nevada. North to British Columbia, east to the 

 Rocky Mts. ]\Iay-July. 



Geog. note. — Holodiscus discolor consists of an assemblage of forms, numerous and varied, 

 without definite geographic segregation, and best represented as a reticulation. The original of 

 the species from Idaho has slightly pubescent rather large thin leaf -blades which are xisiially but 

 not always decurrent on the petiole. In California it tends to become more shortly petiolcd with 

 thicker and smaller leaves which are often more densely pubescent beneath. Specimens from the 

 following stations illustrate its occurrence. Coast Ranges: Eequa, mouth of Klamath River, God- 

 dard 172; Humbug, Siskiyou Co., Butler 494; betw. Willow Creek and Three Creeks, n. Humboldt 

 Co., Jepson 2125 ; Van Duzen River valley, opp. Buck Mt., Tracy 2927 ; Bennett Spr., w. Glenn Co., 

 Heller 11,545 ; Calistoga, Jepson 9973 ; Crockett, Contra Costa Co., Tidestrom; San Mateo, Elmer 

 4274; Los Gates, Heller 7514; Los Angeles, Braunton 462. 



The following forms, here described reluctantly as varieties, are represented by a few typical 

 states and by others less typical, all of which intergrade with each other and with the species. 



Var. franciscanus Jepson comb. n. Leaf-blades thick, white-silky or tomentose beneath, 

 more pubescent above than in the species, commonly rounded or truncate at base, 1 to 2^/4 inches 

 long, not decurrent on the petiole; petioles 1 to 5 lines long; panicles typically large and dense. — 

 Coast Ranges from Del Norte Co. to Santa Cruz Co., and in coastal S. Cal. : Ragged Hill, s. of 

 Crescent City, Jepson 9403 ; Mt. Tamalpais, Jepson 13,860 ; Oakland Hills, Jepson 6815 ; Sara- 

 toga, Santa Cruz Mts., Jepson 5629 ; Pettit Canon, San Luis Obispo, Gondii ; Santa Anita Canon, 

 San Gabriel Mts., Feirson 75. In addition there are numerous intergrades towards the species: 

 leaf -blades densely tomentose but decurrent on petiole (Hupa, n. Humboldt Co., Ghandler 1314, 

 Pacific Grove, Tidestrom, and Santa Cruz Isl., T. Brandegee) ; blades small, pubescent beneath, 

 not decurrent (Mt. Diablo, Jepson 7582) ; leaves smaller, panicles medium (Glenbrook, Lake Co., 

 Jepson 13,859), and also intergrades to the next variety. 



Var. microphyllus Jepson comb. n. Small or dwarfed shrub 1 to 3 (or 4) feet high; leaf- 

 blades spatulate-cuneate or ovoid-flabelliform, decurrent or not decurrent, green and villous above, 

 pale and villous or white-sUky beneath, 4 to 10 lines long, the petioles I/2 to 2 lines long, or the 

 leaves subsessUe; panicles small, narrow {Y2 to 2 or rarely 3 inches long). — Exposed rocky ridges, 

 canon floors and open forests in the mountains, 7000 to 11,000 feet; North Coast Ranges; Sierra 

 Nevada from Butte Co. to Tulare Co. ; White Mts. ; Southern California mountains. 



Locs. — North Coast Ranges: Mt. Sanhcdrin, W. P. Taylor; Little Red Mt., n. Mendocino Co., 

 Jepson 16,506. Sierra Nevada: Colby, Butte Co., B. M. Austin; Summit sta., Nevada Co., Jepson 

 13,857; Glen Alpine, Eldorado Co., Katherine Ghandler; Carson Pass, Jepson 8102; Silver Creek 

 Valley, Alpine Co., Brewer 1974 ; Kennedy Mdw., East Fork Relief Creek, Tuolumne Co., A. L. 

 Grant 185 ; Stubblefield Canon, Tuolumne Co., Jepson 4573 ; Lake Eleanor, Tuolumne Co., Ches- 

 nut 4' Drew; Silver Pass, Fresno Co., A. L. Grant 1528; Kearsarge MUl to Rex Montis Mine, w. 

 Inyo Co., Jepson 892 ; Mt. Whitney, Burton ^ Eyerson 33 ; Coyote Pass, Sawtooth Range, Jepson 

 985 ; Trout Mdws., Tulare Co., Purpus 1854 ; North Fork Middle Tule River, Jepson 4690. White 

 Mts.: SUver Canon, Jepson 7213 (leaves slightly glandular, panicles very ample). S. Cal.: Provi- 

 dence Mts., Mohave Desert, T. Brandegee ; Mt. Pinos, n. Ventura Co., Hall 6673 ; Mt. San Antonio, 

 San Gabriel Mts., Peirson 76 ; Bluff Lake, San Bernardino Mts., Mum 10,573 ; Mt. San Jacinto, 

 A. W. Anthony. 



