1»)G ROSACEAE 



VII. Subgenus Horkelia. Leaves pinnate, the leaflets usually many, V2 to 1% 

 inches Ion?, or often Vr, to Vo inch long; calyx-tube mostly cupulate or campan- 

 ulate; petals -white, rarely pinkish or yellowish; stamens 10, alternately long 

 and short ; filaments dilated, at least the alternate (except one species) ; styles 

 terminal or nearly so ; pistils numerous, or often only 2 to 15. 



35. P. californica Greene. California Honey-dew. Stems stoutish, erect, 

 rather leafy, 1 to 3 feet high ; herbage pubescent and more or less glandular ; leaves 

 pinnate, the basal 3 to 7 (or 15) inches long ; cauline leaves mostly shorter than the 

 basal ; petioles 1 to 4 inches long, or the basal to 8I/2 inches long ; leaflets 9 to 21 

 (or the upper leaves with fewer leaflets), thickish, cuneate-obovate to orbicular 

 or oblong, more or less doubly incised or toothed above the base, 1/2 to 1% inches 

 long ; cyme openly 2 to 4-forked, the branches ending in dense few-flowered clusters 

 and often with solitary flowers in the forks, or sometimes the whole inflorescence 

 eapitately congested ; flowers 4 to 6 lines wide ; calyx-tube cup-shaped, truncatish 

 at base, rusty or purplish, 3 to 5I/2 lines wide; bractlets commonly exceeding the 

 calyx-lobes, 3-toothed, or variably incised, or sometimes entire ; petals oblong or 

 spatulate, commonly erect, about 3 lines long ; stamens 10, the filaments opposite 

 the cah-x-lobes subulate, the others filiform or nearly so; achenes many. 



Wooded slopes or edges of brushy thickets, 5 to 600 feet : Coast Ranges from 

 Humboldt Co. to Monterey Co. May-June. 



Locs.— Blue Lake, Humboldt Co., Tracy 3526; Ft. Bragg, W. C. Mathews 69; Hopland, 

 Jepson 9289 ; Bodega, Crum 753 ; Pt. Reyes, Jepson 1169 ; Silver Spr., Mt. Diablo, M. Bowerman 

 1666 ; Berkeley, Jepson 108p ; Mt. Davidson, San Francisco, Jepson 10,580 ; Niles, Alameda Co., 

 Jepson 118p (varying towards var. frondosa Jepson) ; betw. Salada and Mussel Rock, San Mateo 

 Co., Newlon 250 ; Carmel River, near Carmel, Jepson lllp. 



Note on variation. — Occupying a well-defined range along the coast, Potentilla californica 

 is replaced by the more delicate P. elata at higher altitudes and by P. lindleyi near the coast at 

 the southern limit of its range. It is not known in the inner Coast Range except on Mt. Diablo, 

 Near the ocean in exposed situations the species is often very densely glandular with thickish 

 leaves ; in the shade or inland, usually less glandular and with thinner often more deeply divided 

 leaflets. Its most distinctive characters are the robust habit, the doubly incised leaflets, the 

 large-bracteate few-flowered cymes, the long toothed bractlets, and the large truncate-based 

 calyx-tube which contrasts in color with the sepals. Usually the filaments opposite the calyx-lobes 

 are broader than those alternate, but in some plants this difference is not striking. Occasionally, 

 plants collected near the coast (Pt. Reyes, Jepson 1169) have all filaments nearly equally dUated. 

 The bractlets, typically 3-toothed and exceeding the calyx-lobes, are sometimes entire and of 

 about the same length as the calyx -lobes. The petals usually equal the calyx-lobes (not exceeding 

 them as in P. lindleyi and many other species of the subgenus Horkelia), but here also is con- 

 siderable variation. Var. frondosa Jepson appears to occur only occasionally within the range 

 of the species. Superficially it bears a resemblance to P. glandulosa Lindl. and, as indicated by 

 leaves and inflorescence, is undoubtedly the most primitive Horkelia within our territory. Since 

 so rarely collected one is tempted to consider the form a retrogressive mutation. Intermediates 

 connecting it with the species are uncommon (Crockett, Contra Costa Co., Tidestrom) . 



Var. frondosa Jepson. Leaflets 5 to 9, oblong or oval, toothed or shallowly doubly incised ; 

 calyx-tube usually narrower, 2^^ to 3 lines wide; petals usually shorter than the sepals, IVz to 2 

 lines long. — Sandy or gravelly slopes, 20 to 1500 feet: Contra Costa and Alameda Cos.; Mon- 

 terey Co. 



Locs. — Martinez; Sunol, Alameda Co., Mason; Pt. Pinos, Monterey Co., Michener 4" Bioletti; 

 Jolon grade, Monterey Co., Hall 9969 (less glandular; leaves thin, glabrate). 



Refs. — Potentilla californica Greene, Pitt. 1:100 (1887) ; Jepson, FI. W. Mid. Cal. 282 

 (1901), ed. 2, 209 (1911), Man. 494 (1925). HorJcelia californica C. & S. Linnaea 2:26 (1827), 

 type loc. San Francisco, Chamisso; Rydb. Mem. Dept. Bot. Columbia Univ. 2:126, pi. 57 (1898). 

 Siblaldia californica Spreng. Syst. 4^:341 (1827). Horkelia grandis H. & A. Bot. Beech. 339 

 (1838), type from Cal., Douglas. P. californica var. carmeliana Jepson, Fl. W. Mid. Cal. 282 

 (1901), ed. 2, 209 (1911), type loc. Carmel River, Jepson lllp. Var. frondosa Jepson, Man. 

 494 (1925). P. frondosa Greene, Pitt. 1:300 (1889), type loc. Martinez, Contra Costa Co., 

 Greene. EorTcelia frondosa Rydb. Bull. Torr. Club 25:54 (1898); Mem. Dept. Bot. Columbia 

 Univ. 2:125, pi. 56 (1898). 



36. P. elata Greene. Mountain Honey-dew. Stems erect, 1% to 2 feet high ; 

 herbage glandular, pilose-pubescent; leaves pinnate, the basal with blades 3 to 



