ROSE FAMILY 203 



important are the differences in habit, and in the shape of the calyx-tube. Noteworthy also in 

 var. wilderae is the extreme reduction in the number of pistils. The achenes also are somewhat 

 different in shape and color. The discontinuous distribution of these two closely related forms 

 is no novel occurrence among California species of PotentUla. 



Var. wilderae Jepson. Plants lower (4 to 6 inches high) ; herbage less glandular, more 

 densely silky-pubescent; flowers 2% lines wide; calyx-tube % line high (deeper in proportion to 

 width), truncate at base, strongly 10-nerved in fruit; pistils 1 to 4; achenes smooth, pale. — 

 Montane, 7000 to 8000 feet: San Bernardino Mts. (South Fork Santa Ana Eiver). June-Aug. 



Eefs. — POTENTILLA PARRYi Greene, Pitt. 1:102 (1887) ; Jepson, Man. 494, fig. 490 (1925). 

 Horlcelia parryi Greene, Bull. Cal. Acad. 2:416 (1887), type loc. lone, Amador Co., Edwards, 

 Curran, Parry. E. platypetala Eydb. Bull. Torr. Club 25:55 (1898) ; Mem. Dept. Bot. Columbia 

 Univ. 2:133, pi. 67, figs. 1-5 (1898). Var. wilderae Jepson, Man. 494 (1925). EorTcelia wilderae 

 Parish, Bot. Gaz. 38 : 460 (1904), type loc. trail from Barton Flat to South Fork Santa Ana River, 

 C. M. Wilder 237. P. wilderae M. & J. Bull. So. Cal. Acad. 24:8 (1925). 



47. P. douglasii Greene. Oregon Honey-dew. Steins % to II/2 feet high; 

 herbage minutely hispid to glabrate and usually more or less glandular; leaves 

 mostly basal or sub-basal, the blades 2 to 5 (or 8) inches long, the petioles 1 to 2% 

 inches long; leaflets 9 to 17, cuneate to cuneate-obovate, deeply toothed at apex or 

 above the middle, 8 to 6 or rarely 9 lines long; cymes looselj^ branched with the 

 clusters capitate or subcapitate or the whole cyme capitately congested; flowers 

 21/2 to 3 (or 5) lines wide; calyx-tube bowl-shaped, purplish or pale, often 10- 

 nerved; petals white, cuneate, refuse (rarely truncate or rounded), 1^2 to 2 lines 

 long, exceeding the calyx-lobes; stamens 10; filaments broadly dilated, those op- 

 'posite the petals slightly shorter and narrower; achenes many, smooth or obscurely 

 striate. 



Montane meadows, 4000 to 9500 feet : Trinity and Siskiyou Cos. ; Sierra Nevada 

 from Fresno Co. to Shasta and Modoc Cos. East to Nevada, north to Oregon and 

 Idaho. June-Aug. 



Note on variation. — The plant originally named Horkelia fusca by Lindley (PotentUla 

 douglasii Greene) is the most common and widespread form of a group which, although variable, 

 apparently consists of one complex recognizable over an extensive range in Idaho, Oregon, Nevada 

 and California. The original illustration (Bot. Reg. pi. 1997) is very similar to most of the 

 plants cited below. Watson described var. tenella as a smaller-flowered form, low and slender, 

 ■with leaves more deeply lobed. This is less extreme than var. tenuisecta, and intermediates occur 

 which are very difiicult to separate from the species. Although specimens with deeply divided 

 leaves and open inflorescence present an appearance quite different from that of typical Poten- 

 tUla douglasii, the characters which distinguish var. tenella are not constantly associated. 



Locs. — North Coast Ranges : Big Flat, Salmon Mts., n. Trinity Co., Hall 8633 ; Weed, Siski- 

 you Co., Butler 1868. Sierra Nevada : Nellie Lake, Fresno Co., A. L. Grant 1076 ; Nevada Falls 

 (above), Yosemite, Chesnut 4' Drew (leaflets flnely toothed, intermediate toward var. tenella) ; 

 Fish Camp, Mariposa Co., Jepson 8392 ; betw. Alder Creek and Peregoy Mdws., Yosemite, Jepson 

 4328; Matterhorn Canon, Tuolumne Co., Jepson 3368; Cow Creek, Tuolumne Co., Jepson 6506 

 (cymes open) ; Dorrington, Calaveras Co., Jepson 10,110; Silver Lake, Amador Co., Hansen 292; 

 Carson Pass, Jepson 8104; Mt. Tallac, Jepson 8089; Truckee River, Placer Co., Sonne; Cisco, 

 Placer Co., H. A. Walker 1399 (cymes open); Bear Valley, Nevada Co., Jepson 121p, 122p; 

 Stalkers, Shasta Co., M. S. Baker 298; Eagle Lake (mts. s.), Lassen Co., Baker 4" Nutting; 

 WUlow Creek, Modoc Co., B. M. Austin 110. 



Var. pseudocapitata Jepson comb. n. Stems erect or ascending, 12 to 18 inches high; 

 herbage densely pubescent, scarcely or not at all glandular; basal leaves 8 to 6 inches long; 

 cymes capitate or compact with few branches. — Open places in pine forest, 3500 to 4900 feet: 

 Sierra Co. (Sierraville, Lemmon 89) to Shasta Co. (Elk Flat, Hall 4- Babcock 4119) and Modoc 

 Co. (F orestdale, M. S. Baker). June-Aug. 



Var. tenella Greene. Leaflets more or less deeply incised with linear segments. — Meadows, 

 4300 to 7000 feet: Sierra Nevada, Tehama Co. to Tulare Co. North to Washington. June-Aug. 



Locs. — Cannell Mdws., Tulare Co., Hall $• Babcock 5117; Huntington Lake, Fresno Co., E. 

 Ferguson 372 (intermediate toward the species) ; Little Yosemite, Mariposa Co., Jepson 3150; 

 Jonesville, Butte Co., Heller 11,673; PrattvUle, Plumas Co., Cleveland 159; Warner Valley, 

 Plumas Co., E. M. Austin 388 ; Battle Creek Mdws., Tehama Co., J. Grinnell. 



Var. tenuisecta Jepson comb. n. Leaflets divided to the base into linear segments. — Meadows, 

 4400 to 5100 feet: Plumas Co. (Big Mdws., Jepson 4056) ; Lassen Co. (Martin Spr., Eagle Lake, 

 Brown 4' Wieslander) . North to Washington. June- July. 



