208 ROSACEAE 



100, north temperate regions of all contiuenls, iu)iie in south temperate rejrions, 

 but occurring in mountains of the tropics. (The ancient Latin name.) 



Bibliop. — Lindlcv, J., Eosarum monographia, ed. 2, 1-156, tt. 1-18, — 1830. Crepin, F., 

 prodrome d'une monop;raphio dos roses aiii6ricaincs (Bull. Soc. Bot. Belg. 15:] 2-10'), — ISTT)). 

 Watson, S., History and revision of the roses of N. Am. (Proc. Am. Acad. 20:324-352, — 1885). 

 Willmot, E. A., The genus Kosa (1:1-243,-1910-1911; 2:1-306,-1910-1912; the text descrip- 

 tions accompanied by a large number of unnumbered plates, the greater part beautifully colored). 

 Greene, E. L., Certain western roses (Lflts. 2:254-266,-1912). Eydberg, P. A., Koses of Cali- 

 fornia and Nevada (Bull. Torr. Club 44:65-84, — 1917) ; Eoses of the Columbia River region (I.e. 

 48:159-172,-1921) ; Rosa in X. Am. Fl. 22:483-533,-1918. Tiickholm, G., Zytologische Studien 

 iiber die Gatt. Rosa (Acta Hort. Berg. 7:99^02, figs. 1-56,-1923). Hurst, C. C, Differential 

 polyploidy in the genus Rosa L. (Zeitschr. Indukt. Abstam. u. Vercrbungsl. Siipplenientli. 2:SG6- 

 906,-1928) ; Genetics of the rose (Rose Annual, Rose Soc. London 1929:37-64, pis. 1-11, figs. 

 1-9, — 1929) ; the general conclusion of the author is that there are in the genus 5 differential 

 septets, each representing a distinct set of chromosomes and corresponding characters. There 

 are thus 5 primary species. All other species are polyploids -which may contain any of the 5 

 septets in any combination. Erlanson, Eileen W., Cytological conditions and evidences for 

 hybridity in N. Am. wild roses (Bot. Gaz. 87:443-506, figs. 1-4 and pis. 16-19,-1929) ; Field 

 observations on wild roses of the western U. S. (Mich. Acad. Sci. Arts and Letters. 11 : 117-135, — 

 1930) ; The phenologieal procession in N. Am. wild roses in relation to the polyploid series (I.e. 

 11:137-150) ; American wild roses (American Rose Annual 1932:83-90). 



Calyx -lobes persistent on fruit. 



Leaves commonly resinous-pubescent beneath; Humboldt Co. and northward.. ..1. R. mifkana. 

 Leaves usually not resinous beneath; hips often constricted into a short neck below calyx- 

 lobes. 

 Leaflets usually not glandular-dentate; stipules not glandular-dentate or glandular- 

 ciliolate on the margins (except in 1 or 2 forms of R. pisocarpa) ; plants com- 

 monly 3 to 5 feet high. 

 Stipules narrow, with the free tips commonly lanceolate. 



Spines usually stout, often curved; calyx-lobes 1 to 2 lines wide, more fre- 

 quently non-glandular than glandular; herbage mostly pubescent; 



cismontane Cal 2. Jk. calif ornica. 



Spines slender, straight; calyx-lobes narrow (1 to 1^/^ lines wide), usually not 

 glandular ; herbage glabrate or nearly ; mostly desert and Great Basin 



areas 3. B. woodsii. 



Stipules usually strongly enlarged upwards and sagittate in outline; herbage sub- 

 glabrous or somewhat puberulent, especially on the leaves beneath ; calyx- 

 lobes relatively short and broad, usually glandular ; Lake Co. to Siskiyou 



Co 4. E. pisocarpa. 



Leaflets usually glandular-dentate ; stipules usually glandular-eiliolate or glandular- 

 dentate on the margins ; plants % to 3 feet high. 



Calyx-tube glandular-hispid; Coast Ranges and Sierra Nevada 5. B. spithamea. 



Calyx-tube glabrous; Sierra Nevada chiefly 6. B. pinetorum. 



Calyx -lobes deciduous from fruit; Coast Ranges and Sierra Nevada 7. B. gymnocarpa. 



1. R. nutkana Presl. Nootka Rose. Stout, 2 to 5 feet high ; prickles stout 

 (rarely slender) , usually straight, or the stem sometimes unarmed ; leaves resinous- 

 pubescent beneath or merely puberulent; stipules subequal in width from base to 

 apex, mostly glandular-ciliate ; flowers solitary or 2 to 4 together, the pedicels often 

 prickly and glandular; calyx-tube glabrous; sepals prolonged into foliaceous, ser- 

 rate or laciniate appendages; petals obcordate, % to 1% inches long; hips globose 

 or depressed-globose, 5 to 7 lines in diameter. 



Valley flats or hillslopes, 5 to 1500 feet : Humboldt and Siskiyou Cos. East to 

 Utah, north to Alaska. May-June. 



Loes. — Eureka, M. S. BaTcer 11; Bald Mt., n. Humboldt Co., Tracy 6345; Trinidad, n. Hum- 

 boldt Co., Geo. Parrish. 



Var. hispida Fer. Calyx-tube w^ith gland-tipped bristles. — Humboldt Co.: Eureka, Tracy 

 6564. North to British Columbia, east to Utah and Montana. June-July. 



Eefs. — Rosa nutkana Presl, Epim. Bot. 203 (1849), type loc. Nootka Sound, B. C, Eaenlce; 

 Jepson, Man. 498 (1925). B. fraxinifolia Hook. Fl. Bor. Am. 1:199 (1834) ; not R. fraxinifolia 

 Borkh. (1790). B. lyalli<ina Crepin, Bull. Soc. Bot. Belg. 15:39 (1876), as synonym. B. caryo- 

 carpa Dougl. ; Crepin, I.e., as synonym. B. woodsii Regel, Acta Hort. Petrop. 5:299 (1877) ; not 



