580 ONAGRACEAE 



at base; capsules teretisli. ratlier strontrlv curved, obscurely 2-ribbed on the sides, 

 4^2 to 10 lines lonp. 



~l)ry valleys and hill slopes. 750 to 1000 feet: upper Salinas River watershed 

 in Monterey and San Luis Obispo Cos., ea.st of the Santa Lucia Mts. May-July. 



Loca. — San Mifjuelito Rancho, Jcpsun 1G24 ; Jolon, Monterey Co., K. Brandegee ; Paso Robles, 

 Benj. Cobb 'l^^■, Templeton, Davy 7595; Atascadero, C. L. Ilitchcoclc 8; Santa Margarita, K. 

 Brandegee : La Panza, San Juan River, Summers. 



Var. margaritae Iltck. Petals rose-red, with yellow base, 6 to 12 lines long; filaments un- 

 equal. — Dry slopes, 100 to 1000 feet: Santa Margarita Valley; San Luis Obispo Co. west of the 

 Santa Lucia Mts. 



Locs. — Santa Margarita Valley, lihoda Meed; Cavitas Canon, San Luis Obispo, BJioda Reed; 

 Price Canon, sc. of Edna, C. L. Ilitchcoclc 36; betw. Arroyo Grande and Pismo, Ehoda Reed. 



Var. luteola Iltck. Petals 5 to 10 lines long, cream-color with large crimson central spot; 

 filaments unoqual.^ — Grassy hillsides, San Luis Obispo Co.: McGinnis, Palmer 143; betw. Atasca- 

 dero and Morro, C. L. Ilitchcoclc 40. 



Befs.— GODETIA PARVIFLORA Jepson, Univ. Cal. Publ. Bot. 2:339 (1907), Fl. W. Mid. Cal. ed. 

 2, 280 (1911), Man. 677 (1925). Oenothera viminea var. parviflora II. & A. Bot. Beech. 342 

 (1840), tvpe from Cal., Douglas, the exact locality not known (as generally in Douglas plants 

 from Cal."), but doubtless Monterey Co. 0. tenella Wats. Proc. Am. Acad. 8:598 (1873), in 

 part; not O. tenella Cav. (1797). G. quadrivitlnera var. tenella Jepson, Fl. W. Mid. Cal. 334 

 (1901), in part. Var. margarit.\e Htck. Bot. Gaz. 89:350 (1930). G. viminea var. margaritae 

 Jepson, Univ. Cal. Publ. Bot. 2:339 (1907), type loc. Santa Margarita Valley, San Luis Obispo 

 Co., Summers; Man. 677 (1925). Var. luteola Iltck. Bot. Gaz. 89:349 (1930), type loc. "top 

 of grade between Atascadero and Morro Beach, San Luis Obispo Co.", C. L. Ilitchcoclc 40. 



5. G. quadrivulnera Spach. Four-spot. Stem erect, simple or with simple 

 branches from below the middle, % to 1% feet high; herbage pubescent or pubeni- 

 lent; blades of lowest leaves obovate or oblong, 5 to 11 lines long, the upper narrowly 

 or broadly linear, the uppermost lanceolate and somewhat conduplicate, % to 1% 

 inches long; flowers discrete or remote; calyx-lobes usually distinct and reflexed 

 (in the earliest flowers sometimes united and turned to one side) ; petals lilac or 

 pale crimson, usually with a spot at the top, 2^2 to 6 (or 7) lines long; ovary canes- 

 cent to densely villous; stigmas purple, short-oblong; capsules sharply 4-sided, 

 8-ribbed or the ribs sometimes obscure, shortly beaked or sometimes bluntish, ^ 

 to 1 inch long, sessile. 



Dry hillsides, 20 to 4400 feet : coastal Southern California; Coast Ranges from 

 San Luis Obispo Co. to Humboldt Co.; Sacramento Valley; Sierra Nevada from 

 Tulare Co. to Plumas Co. North to Washington. May-July. 



Note on variation. — Of Californian species of Godetia, Godetia quadrivulnera is the most 

 widely distributed and the most variable. Variability in shape, size and color of the petals is a 

 striking feature. In one colony a few yards square, where the individuals closely match each 

 other and appear to be of one genetic constitution, a wide range of petal form may occur: petals 

 truncate and erosulate, petals truncate with a notch, petals truncate and erosulate with a central 

 spear point, petals broadly obovate, petals narrowly obovate, and petals extremely narrow with a 

 spear point set in the cleft at apex, petals linear and 3-lobed at apex (Owens Creek, Mariposa Co., 

 Jepson 12,760, 12,761). Pubescence is not only variable as from one locality to another, but the 

 degree of it changes from the juvenile to the senile stages. Plants bearing flowers as large or 

 nearly as large as those of Godetia purpurea occur and when the inflorescence is variously con- 

 gested represent intergrades to that species, as represented by Godetia quadrivulnera var. elmeri 

 Jepson. 



Locs. — S. Cal. : Mesa Grande, San Diego Co., E. Ferguson 28 ; Ramona, San Diego Co., K. 

 Brandegee; Lakeside, San Diego Co., Rail 7438; Fullers Mill, San Jacinto Mts., Hall 2262; Red- 

 lands, Condit ; San Bernardino foothills. Parish; San Antonio Caiion, San Gabriel Mts., Peirson 

 467; Griffith Park, Los Angeles, Braunton 401; Santa Barbara, M. S. Baker. Coast Ranges: 

 Arroyo Grande, Summers; Lopez Canon, San Luis Obispo, Rhoda Reed; Waltham Creek, San 

 Carlos Range, Jepson 2663; Santa Margarita, Rhoda Reed; Paso Robles, Barber; San Miguelito 

 Rancho, Santa Lucia Mts., Jepson 1626; Ventana Cone, Monterey Co., Davy 7394; Seaside, 

 Monterey Co., Heller 6754; Gilroy (hiUs w.), Jepson 14,338; Eva sta., Santa Cruz Mts., Jepson 

 14,340; Belmont, San Mateo Co., Davy 792; Alameda, B dander ; Mt. Tamalpais, Jepson 14,348; 

 Hoods Peak, Sonoma Co., Bioletti; St. Helena, Jepson 14,344; Houghs Sprs., ne. Lake Co., Jep- 

 son 9010; Bennett Spr., Glenn Co., Heller 11,540; Round Valley, Mendocino Co., P. E. Goddard 

 606; Cummings, n. Mendocino Co., Jepson 1873; Rush Creek, Trinity Co., H. S. Yates 405; 

 Kneeland Prairie, Humboldt Co., Tracy 3831; betw. Orleans Bar and Sommes Bar, Klamath 



