582 ONAGRACEAE 



but slijjlitly free; calyx-tube 2V2 to 3Vo lines long, the lobes reflexed, distinct; 

 petals broadly euneate, truncatish and eroded at apex, 7 to 12 lines long, light 

 crimson, usiuilly with a wedge-sliapcd darker spot in middle at apex; stigmas 

 oval, purple; ovary shaggy-villous or eanescent; capsules 4-sided, sessile, stoutish 

 and mostly sluu-t. 4 to 7 (or 11) lines long, shaggy or merely pubescent, 8-ribbed; 

 seeds faintly granulate on sides. 



Plains or valley Hats, often on margins of former pools, 50 to 1500 feet : Sierra 

 Nevada foothills from Butte Co. to Mariposa Co.; Great Valley; Coast Ranges 

 from IMendocino Co. to Santa Cruz Co. ; south to Santa Barbara Co. Mar.-May, 



Geog. note. — Godctia purpurea is closely related to G. quadrivulnera. The two seem to be 

 connected by many different phases or intergrades. The natural type of G. purpurea, the ex- 

 treme phase furthest removed from G. quadrivulnera and therefore best marked, is the plant of 

 the Sacramento Valley plains which grows in "hog-wallows" or on the clay edges of former winter 

 pools or ponds. It is low, simple-stemmed or nearly so, the few flowers congested at apex, with 

 the capsules markedly short (4 to 6 lines long) and strongly ridged, sometimes almost as if nar- 

 rowly winged. In the Coast Range phases of it, the flowers are frequently smaller and the capsules 

 often tend to elongate and be more hairy. Plants with looser inflorescence, smaller flowers and 

 longer capsules represent intergrades to G. quadrivulnera. 



The contrasts between Godetia purpurea and G. viminea may well be emphasized. The stems 

 in G. purpurea are usually simple and they are also very leafy with broadish leaves. The short 

 or condensed terminal inflorescence is very leafy, the leaves usually broader and longer than the 

 cauline leaves and often nearly concealing the flowers. The stems in Godetia viminea and in its 

 variety williamsonii are tall and branching and only moderately leafy. The leaves are narrower 

 than in G. purpurea and the inflorescence is looser, much less leafy and with the leaves reduced. 

 The calyx-tube in G. viminea is longer than in G. purpurea and is nearly always whitish. In addi- 

 tion to the differences in morphological characters, the geographic areas of distribution in these 

 two species are mutually exclusive. Godetia purpurea is a plant of the Coast Range valley floors 

 or of the Great Valley plains at low altitudes, about 1 to 1500 feet. On the other hand G. vimi- 

 nea, with its varieties, inhabits the Sierra Nevada at middle altitudes and is also found in its 

 foothills but does not occur on the plains or valley floors to the west. The following are cited as 

 G. purpurea. 



Locs. — Sierra Nevada foothills: Clear Creek, Butte Co., H. E. Brown 212; Ruth Pierce Mine, 

 Mariposa Co., Jepson 10,713. Great Valley: Willows, Glenn Co., Jepson 14,370; College City, 

 Colusa Co., Alice King ; Lincoln, Placer Co., Hall 10,291 ; Browns Valley, n^Y. Solano Co., Jepson 

 14,369; Elmira, Solano Co., K. Brandegee; Oakdale, Stanislaus Co., Jepson 14,368; Sanger, 

 Fresno Co., Gondii. Coast Ranges: Potter Valley, Mendocino Co., Pur pus ; Cache Creek Canon, 

 Yolo Co., C. F. Baker 5065; Sebastopol, E. Ferguson 226; Tiburon, E. A. Walker 1736; Oakland 

 Hills, Bolander; Pescadero, Santa Cruz Co., Guirado 690. Santa Barbara Co.: Santa Maria, 

 Eastwood. 



Var. arnottii Jepson comb. n. Herbage glabrous; capsules short (4 to 5 lines), glabrous. — 

 Margins of winter pools on the sandy-clay plains and valley floors, 1 to 500 feet: Solano Co. 

 (Little Oak, Vacaville, Jepson 14,334; Elmira, C. F. Baker). Also collected by Douglas, doubt- 

 less in the South Coast Range valleys, his specimen at Kew labeled by William Hooker as Oeno- 

 thera arnottii T. & G. All the specimens just cited represent Oenothera arnottii T. & G. Fl. 1 :503 

 as to the diagnosis, but are different from the Douglas plant preserved in the Gray Herbarium, 

 which is G. purpurea G. Don. 



Refs. — Godetia purpurea G. Don; Sweet. Hort. Brit. ed. 3, 237 (1839) ; Jepson, Univ. CaL 

 Publ.Bot. 2:344 (1907), Fl. W. Mid. Cal. ed. 2, 280 (1911), Man. 679 (1925). Oenothera purpurea 

 Curt. Bot. Mag. t. 352 (1796), type collected by Menzies, on the "western coast of North America", 

 probably between San Francisco and Monterey (typ. vidi). Clarkia purpurea Nels. & Mcbr. Bot. 

 Gaz. 65:64 (1918), in part. G. lepida Lindl. Bot. Reg. t. 1849 (1836), type from Cal., Bonglas; 

 Jepson, Fl. W. Mid. Cal. 335 (1901). G. purpurea var. lacunarum Jepson, Univ. Cal. Publ. Bot. 

 2:346 (1907), type loc. Oakdale, Stanislaus Co., Jepson 14,374; Man. 679 (1925), stem strictly 

 branched. G. sparsifolia Jepson, Univ. Cal. Publ. Bot. 2:347 (1907), type loc. Tracy, B. Cobb; 

 Fl. W. Mid. Cal. ed. 2, 281 (1911) ; Man. 679 (1925). Var. arnottii Jepson. Oenothera arnottii 

 T. & G. Fl. 1:503 (1840), type from Cal., Douglas. 0. viminea var. ("erecta glabriuscula glau- 

 cescens") H. & A. Bot. Beech. 342 (1840), California, Douglas. G. arnotUi Jepson, Univ. Cal. 

 Publ. Bot. 2:346 (1907), Fl. W. Mid. Cal. ed. 2, 281 (1911), Man. 679 (1925). 



7. G. arcuata Jepson. Kellogg Godetla.. Stem slender, sparingly branched, 

 more or less falsely dichotomous on account of the repeated divergence of axis and 

 branch, 7 to 20 inches high; leaf -blades linear, entire, obtusish, I14 to 2 inches 

 long, the fascicled and upper leaves narrow (I/2 to 1 line broad), commonly con- 

 duplicate and appearing much narrower, more or less arcuate-curving, with long 



